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С ъстояние на морските екосистеми пред българското черноморско крайбрежие. Цонка Консулова Институт по Океанология към БАН - Варна. УВОД Черно море е уникален морски басейн , със следните собености :. От всички континентални морета, то е най-изолирано от световния океан. - PowerPoint PPT Presentation

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    TEMP

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    DEPTH

    CTD - PROFILES

    Sheet1

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    3333

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    10101010

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    12121212

    13131313

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    19191919

    20202020

    21212121

    22222222

    23232323

    24242424

    25252525

    26262626

    27272727

    28282828

    29292929

    30303030

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    34343434

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    DEPTH

    CTD - PROFILES

  • , .

  • , ( ) .

  • , .

  • .

  • 80- ( ) (, ) -

  • Mnemiopsis leidyi :. ., - Mnemiopsis leidyi, 80- .

    90- Cladocera Copepod 45%.

    Mn. leidyi 70% , .

  • 80- - , -, - -, - .

    90- . (Kamburska et al., 2003)

    80- ..() 1989-1991 . 1993, 1994-1997 ., 1977-1988 (B). 80- - M.Leidyii - , .

    Chart1

    7074422.53

    415342.565

    20588.38

    191158.485

    543372.91

    3698.5240.46

    297350.465

    2066.44

    20117.06

    477530.2

    12561937.51

    4163.5366.08

    20322.56

    20875.5482.595

    7954626.52

    705764.61

    13366.552.155

    9169.5110.58

    22474.5102.095

    10733180.23

    412673.28

    2030.2

    12342460.2

    667146.5

    5452.64

    386240.03

    9931.5220

    2098318.815

    510216.83

    14702390.85

    36955181.24

    20225.7

    year

    [mg.m-3]

    [ind.m-3]

    A

    N.scintillans [ind.m-3]

    zooplankton biomass [mg.m-3]

    Sheet1

    Sheet1

    7074422.53

    415342.565

    20588.38

    191158.485

    543372.91

    3698.5240.46

    297350.465

    2066.44

    20117.06

    477530.2

    12561937.51

    4163.5366.08

    20322.56

    20875.5482.595

    7954626.52

    705764.61

    13366.552.155

    9169.5110.58

    22474.5102.095

    10733180.23

    412673.28

    2030.2

    12342460.2

    667146.5

    5452.64

    386240.03

    9931.5220

    2098318.815

    510216.83

    14702390.85

    36955181.24

    20225.7

    year

    [mg.m-3]

    [ind.m-3]

    A

    N.scintillans [ind.m-3]

    zooplankton biomass [mg.m-3]

    0026

    00370

    001890

    001530

    00580

    00190

    0085

    00235

    00490

    00200

    00305

    00

    00

    00

    00

    00

    00

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    year

    [g.m-2]

    B

    Catches

    Total biomass

    M.leidyi wet weight [g.m-2]

    anchovy biomass in thousand tons

    2.769154.14259.731983

    15.009166666724.055.921984

    70.123.3251.351985

    12.405246.252.073.52

    1.523.4751.63.2

    3.72.6257.252.9

    4.87595.533333333319891.5

    5.3751.425199010.3

    15.053.82.390

    6.841.45.63.7

    9.50753.5207.225

    4.706252.9525199413.7

    2.64.025183.2

    3.64166666677.719961996

    1.85105.2519971.1

    2.044166666776.3519981998

    year

    N [1x106 cells/l]

    C

    Bacillariophyceae

    Dinophyceae

    Euglenophyceae

    Chrysophyceae

    Chart5

    2.769154.14259.731983

    15.009166666724.055.921984

    70.123.3251.351985

    12.405246.252.073.52

    1.523.4751.63.2

    3.72.6257.252.9

    4.87595.533333333319891.5

    5.3751.425199010.3

    15.053.82.390

    6.841.45.63.7

    9.50753.5207.225

    4.706252.9525199413.7

    2.64.025183.2

    3.64166666677.719961996

    1.85105.2519971.1

    2.044166666776.3519981998

    year

    N [1x106 cells/l]

    C

    Bacillariophyceae

    Dinophyceae

    Euglenophyceae

    Chrysophyceae

    Sheet1

    Sheet1

    7074422.53

    415342.565

    20588.38

    191158.485

    543372.91

    3698.5240.46

    297350.465

    2066.44

    20117.06

    477530.2

    12561937.51

    4163.5366.08

    20322.56

    20875.5482.595

    7954626.52

    705764.61

    13366.552.155

    9169.5110.58

    22474.5102.095

    10733180.23

    412673.28

    2030.2

    12342460.2

    667146.5

    5452.64

    386240.03

    9931.5220

    2098318.815

    510216.83

    14702390.85

    36955181.24

    20225.7

    year

    [mg.m-3]

    [ind.m-3]

    A

    N.scintillans [ind.m-3]

    zooplankton biomass [mg.m-3]

    87.9616.626

    123.4706370

    152.7768.71890

    152.57511530

    225.5707.8580

    233.1642.3190

    364.4647.885

    418.6602.8235

    415.2717.4490

    368.9670.5200

    316.6779.3305

    86.41006

    197.91421.9

    279.91271.5

    373.7821122

    273.239986.7

    213.781003.9

    863.7

    1043.9

    1060

    826.9

    446.1

    365.6

    243.3

    430.5

    592.1

    637.1

    year

    [g.m-2]

    B

    Catches

    Total biomass

    M.leidyi wet weight [g.m-2]

    anchovy biomass in thousand tons

    2.769154.14259.731983

    15.009166666724.055.921984

    70.123.3251.351985

    12.405246.252.073.52

    1.523.4751.63.2

    3.72.6257.252.9

    4.87595.533333333319891.5

    5.3751.425199010.3

    15.053.82.390

    6.841.45.63.7

    9.50753.5207.225

    4.706252.9525199413.7

    2.64.025183.2

    3.64166666677.719961996

    1.85105.2519971.1

    2.044166666776.3519981998

    year

    N [1x106 cells/l]

    C

    Bacillariophyceae

    Dinophyceae

    Euglenophyceae

    Chrysophyceae

  • :

    Broe ovata, Mn. laidyi .

    , .

    (Konsulov, Kambursk, 1998).

    :. .

  • 5. M. leidyi 1995 . (: Yu. Zaitsev ).

  • Chart2

    2521

    2818

    4018

    76.513

    8416

    9312

    10312

    1181

    1401

    Depth (m)

    S

    Data

    , 1998

    Location43o10'2N 28o11'4E43o11'0N 28o14'8E43o10'5N 28o29'2E43o41'1N 28o29'2E43o09'2N 28o32'3E43o09'3N 28o39'0E43o09'6N 28o41'1E43o09'7N 28o41'8E43o09'9N 28o42'9E

    Depth (m)25284076.58493103118140

    AbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomassAbundanceBiomass

    Total Abundance/Biomass1832383.66082176273.05003236712.76085883.2648102060.1202110835.2980415699.2334120.001320.013

    S21211818181813131616121212121111

    H'3.151.652.902.372.820.302.842.402.620.831.780.870.790.220.000.000.000.00

    J0.720.380.700.570.680.070.770.650.650.210.500.240.220.060.000.000.000.00

    Actinothoe clavata321.68804410.816480.2904-0.101994826-0.0344426458-0.1138021019-0.184513517-0.0214091369-0.0045881327000000000000

    Pachycerianthis solitarius43.1460000000000-0.029291488-0.3108741287000000

    Micrura fasciolata80.016440.022400000000-0.054857674-0.003229779500-0.0096448315-0.00273429810000

    000000000000000000

    Nemertini g.sp.40.0840120.0288161.0976160.0224200.0252120.0024-0.0192995716-0.0026617287-0.0413740828-0.0013934125-0.0378738981-0.0143874228-0.1414876829-0.0493133616-0.1112240263-0.004703066500-0.0243580869-0.00037089540000

    000000000000000000

    Aonides paucibranchiata560.35761480.50562281.181600000000-0.2298744283-0.043976322-0.3879374932-0.0877394068-0.2297604675-0.07611148620000

    Aricidea claudiae80.0020000000000-0.0513627593-0.0007993257000000

    Capitelides giardi320.0204000000000000-0.0540595183-0.00251790260000

    Capitomastus minimus1320.0582240.2064360.0228240.02800-0.2452596835-0.0025916419-0.2666846654-0.0034036326-0.2467192251-0.0500156288-0.1272797867-0.005154836500000000

    Exogone gemmifera200.0124320.006400000000-0.1112240263-0.0025252219-0.1476893044-0.0022535853000000

    Harmothoe reticulata320.571240.0160000-0.0658590447-0.0082424705000000-0.0096448315-0.00203133830000

    Melinna palmata48415.43847480.43281614.3744120.0476-0.5073388791-0.1865205693-0.5295703811-0.0147199877-0.5011920795-0.1135786135-0.1145859152-0.08893575210000000000

    Nephthys hombergii1964.30804928.264047.23041120.5872-0.3449786097-0.0727243737-0.4849791274-0.1526724966-0.3747592403-0.067187163-0.4556795091-0.44516324650000000000

    Nereis diversicolor000000000000000000

    Nereis succinea40.0360-0.0192995716-0.00125544150000000000000000

    Pectinaria koreni49.0196161.259200-0.0167048949-0.1625198762-0.0378738981-0.0161556167000000000000

    Polydora ciliata760.046440.0004480.0192-0.1904677931-0.001573845-0.0167048949-0.0000283915-0.0901117438-0.000408912000000000000

    Prionospio cirrifera2840.05762360.0676960.0096-0.4169245054-0.0019069057-0.3475815583-0.0029658969-0.1505572329-0.0002179248000000000000

    Phyllodoce maculata400.1908840.177280.0144880.1484000000-0.2637921258-0.2394288735-0.29663826-0.0247770502-0.0513627593-0.0045932899-0.1177613696-0.01403521720000

    Leiochone clypeata40.3376000000-0.0489773629-0.33851309990000000000

    Notomastus profundus80.018120.0468120.001320.0130000000000-0.0513627593-0.0055774471-0.0243580869-0.00521139810000

    Oridia armani1560.0121800.0412200.013600-0.272574199-0.000636096400-0.5228018108-0.0796068863-0.1112240263-0.002739451500000000

    Sphaerosyllis bul-bosa40.000400-0.0167048949-0.000028391500000000000000

    Spio filicornis160.004200.0128000000-0.1414876829-0.0118511146-0.1112240263-0.002596928700000000

    Syllis hialina000000000000000000

    Terebellides stroemi401.239640.05160.7984000000-0.2637921258-0.5304671664-0.0313504056-0.0085093746-0.0882850854-0.123642093000000

    000000000000000000

    Amphipoda g. sp.160.0160-0.0597310375-0.0006067640000000000000000

    Ampelisca diadema960.4524360.2184720.1848480.368160.0048-0.2229296744-0.0114709524-0.0979004422-0.0082288741-0.1221523808-0.0030887870000-0.1961926209-0.0686388794-0.0308796148-0.00069341990000

    Apseudopsis ostroumovi640.04940.021200000000-0.250626098-0.008362942300-0.0096448315-0.0026047880000

    Balanus improvisus80.4160160.3872-0.034232331-0.010679218100-0.0378738981-0.0058920453000000000000

    Caprella acantifera variabilis40.00480000-0.0119406624-0.0001156968000000000000

    Coremapus versiculatus000000000000000000

    Corophium runcicorne160.0024720.0336320.014-0.0597310375-0.000108135800-0.1221523808-0.000677536800-0.156685598-0.002810285100000000

    Eudorella truncatula80.00316000000-0.0843492836-0.00969154360000000000

    Iphinoe elisae80.0040200.0068-0.034232331-0.0001725427-0.0621832238-0.000380861500000000000000

    Microdeutopus gryllotalpa40.00200000000-0.0313504056-0.000494860700000000

    Phtisica marina40.0004160.01923400.3352-0.0192995716-0.0000207177-0.0521136974-0.0009700744-0.3415345133-0.005198604000000000000

    000000000000000000

    Cardium edule40.0284000000-0.0489773629-0.05954384350000000000

    Cardium paucicostatum161.1680-0.0597310375-0.02544974110000000000000000

    Chamelea gallina80.5240-0.034232331-0.01299692230000000000000000

    Modiola phaseolina5329.802474430.1328368096.78700000000-0.4897899733-0.4266349781-0.3858213141-0.1948513505-0.1553904686-0.0351246850000

    Mya arenaria3239.2160169.2224160.0528-0.101994826-0.3363207143-0.0521136974-0.1650905558-0.0378738981-0.0010163961000000000000

    Mytilus galloprovincialis1024686.393610449.09080000-0.5252891823-0.052370141100-0.3358500262-0.238755178500000000

    Pitar rudis606.89202045.5936-0.1615386417-0.1041677633-0.0621832238-0.431183363100000000000000

    Spisula subtruncata380262.7192148119.046-0.4707163255-0.3740962191-0.2637616739-0.522156993100000000000000

    Syndesmya alba40.0052000000-0.0489773629-0.01480356840000000000

    Syndesmya frafilis40.0040160.032-0.0192995716-0.000172542700-0.0378738981-0.0006484333000000000000

    Tapes lineatus8441.04405651.6948-0.2038965364-0.3449661654-0.1358851303-0.454579910700000000000000

    000000000000000000

    Nassa reticulata209.54403218.5536-0.071149298-0.1325672795-0.0895215124-0.26360273400000000000000

    000000000000000000

    Phoronis euxinicola160.27841160.734880.00520000-0.0378738981-0.0044223206-0.4619663207-0.4842504332-0.054857674-0.001167405700000000

    Leptosynapta inchaerens200.4636320.094400000000-0.1112240263-0.0541236967-0.1476893044-0.0228567487000000

    Amphiura stepanovi40.0164560.48880000000000-0.029291488-0.0051440724-0.0837254574-0.03775811310000

    Molgula euprocta560.1956200.49360000000000-0.2176513409-0.041535693-0.0370502976-0.03805877070000

    3.15301830691.65488118792.90091841982.36826307512.82088565160.30159984962.84359377052.40158451852.61528046130.83056137861.78393771730.86850602070.78627786220.21725231270000

    Sheet1

    Sheet1

    2521

    2818

    4018

    76.513

    8416

    9312

    10312

    1181

    1401

    Depth (m)

    S

    Depth (m)

    Number of species

    Indeces

    Depth (m)SH'JTotal Abundance

    25213.150.721832

    28182.900.702176

    40182.820.683236

    76.5132.840.77588

    84162.620.651020

    93121.780.501108

    103120.790.224156

    11810.0012

    14010.0032

    Depth (m)SH'JTotal BiomassStationDepthBiomassSH'(B)J(B)

    25211.650.38383.66130125387.883351.780.35

    28182.370.57273.05030227287.033381.970.38

    40180.300.07712.7613033054.669322.260.45

    76.5132.400.653.2653047816.027212.500.57

    84160.830.2160.12030595106.930180.39007289790.0935443438

    93120.870.2435.298

    103120.220.0699.233

    11810.000.001

    14010.000.013

    November 1988September 2000

    Depth (m)AbundanceSH'(A)J(A)DepthAbundanceSH'(A)J(A)

    251832213.150.722525910352.130.42

    282176182.900.702712490383.180.61

    403236182.820.683010100323.160.63

    76.5588132.840.77782220213.470.79

    931108121.780.50952150182.670.64

    Average52.5178816.42.700.67511057428.82.920.62

    Depth (m)BiomassSH'(A)J(A)DepthBiomassSH'(A)J(A)

    25383.661211.650.3825387.883351.780.35

    28273.050182.370.5727287.033381.970.38

    40712.761180.300.073054.669322.260.45

    76.53.265132.400.657816.027212.500.57

    9335.298120.870.2495106.930180.390.09

    Average52.5281.60716.41.520.3851170.50828.81.780.37

    t-Test: Paired Two Sample for Means

    Variable 1Variable 2

    Mean178810574

    Variance103789694960430

    Observations55

    Pearson Correlation0.4151257287

    Hypothesized Mean Difference0

    df4

    t Stat-2.0971919037

    P(T

  • 60- : - . o, 1960). 142 90 45

  • .

    - , - , , 1954-1957.

    . 11. - ( ) .( ., 2004)

  • 10-

    () - () , , .

    (a) (b) - b( ., 2004)

  • , . - :

    .

    - .

    , - .

  • Rapana : , 25-50 . Rapana , 9-10 , - . 93.7% .

  • ( t) ( ).

    Chart1

    42746

    72971

    63618

    88949

    1191115

    668115

    642112

    Meat yield in tons

    Number of trawlers

    Years

    Tons

    Sheet2

    1996427

    1997729

    1998636

    1999889

    20001191

    R. thomasiana total quantity meat exported from Bulgaria during the period 1996-2000

    Sheet2

    0

    0

    0

    0

    0

    Size comp

    Size comp

    0

    0.3344481605

    1.0033444816

    0.6688963211

    1.0033444816

    0.6688963211

    1.0033444816

    1.0033444816

    1.6722408027

    1.0033444816

    2.6755852843

    3.0100334448

    4.0133779264

    3.3444816054

    3.6789297659

    4.0133779264

    4.347826087

    3.0100334448

    3.3444816054

    4.6822742475

    5.016722408

    3.3444816054

    3.6789297659

    0.6688963211

    3.6789297659

    2.0066889632

    2.3411371237

    2.0066889632

    3.3444816054

    2.6755852843

    1.3377926421

    2.0066889632

    0.3344481605

    0.3344481605

    0.3344481605

    1.3377926421

    4.347826087

    10.3678929766

    6.3545150502

    mm

    %

    Sheet1

    0

    0.826446281

    4.958677686

    3.305785124

    1.652892562

    3.305785124

    1.652892562

    0

    12.3966942149

    4.958677686

    4.958677686

    2.479338843

    2.479338843

    0.826446281

    2.479338843

    0

    4.132231405

    0.826446281

    0

    0.826446281

    0.826446281

    0

    0.826446281

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0.826446281

    6.6115702479

    9.0909090909

    19.8347107438

    9.9173553719

    mm

    %

    0

    0.1414427157

    0.1414427157

    0

    0.1414427157

    0.1414427157

    0.1414427157

    0.2828854314

    0

    0.2828854314

    0.1414427157

    0.5657708628

    0

    0.4243281471

    0.7072135785

    0.8486562942

    1.2729844413

    3.1117397454

    4.5261669024

    10.6082036775

    10.1838755304

    13.1541725601

    14.4271570014

    10.4667609618

    9.1937765205

    4.3847241867

    1.9801980198

    1.5558698727

    1.1315417256

    0.9900990099

    1.8387553041

    2.2630834512

    2.5459688826

    2.4045261669

    0

    mm

    %

    ? %

    19962319962024815423.876.21996

    19974219972789218633.166.91997

    1998251998120754562.537.51998

    19998419992121337962.737.31999

    2000122200041526714864.335.72000

    2001215200141626615063.936.12001

    % ( 1) (.2)

    Number of trawling for Rapana thomasiana vessels along the Bulgarian Black sea

    coast during the period 1996 - 2001

    .

    19962024272021996

    19972787292781997

    19981206361201998

    19992128892121999

    200041511914152000

    20014166684162001

    1996199719981999200020012002

    Number of licences202278120212415416185

    Meat yield t.4277296368891191668642Data for Mean Yield in tons of Rapana and number of licences during the period 1996-2002

    199646

    199771Number

    199818YearsMeat yield in tonsof trawlers

    199949199642746

    2000115199772971

    2001115199863618

    199988949

    20001191115

    2001668115

    2002642112

    0

    0

    0

    0

    0

    0

    00

    00

    00

    00

    00

    00

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    00

    00

    00

    00

    00

    00

    %

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    00

    00

    00

    00

    00

    00

    Meat yield t.

    Number of licences

    Meat yield in t

    Number of licences

    00

    00

    00

    00

    00

    00

    00

    &A

    Page &P

    Number of licences

    Meat yield t.

    Number of licences

    Mean yield in t

    00

    00

    00

    00

    00

    00

    00

    Meat yield in tons

    Number of trawlers

    Years

    Tons

    00

    00

    00

    00

    00

    00

    00

    Meat yield in tons

    Number of trawlers

    Years

    Tons

  • , , , a 2-3 .

  • A

    , 2001 2002 ( , ). . .

    Upogebia pusilla

    Upogebia pusilla

    (Erifia verrucosa)

    . Gobiidae

  • (650 ) 18 . 1999 .

  • - .

  • (MP) (SP) (PS) (TS).

  • (PS) (S) 2 - 4 - PS. PS S. () 22.8 %, (b) 4 - - 6.3%. () (b)b

  • - (), (b) ( c ) (2001 ) , 4, 9 6 2000 . - Crustacea (13 93 ), Mollusca 6 .199920002001

    Chart1

    4113

    913104

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    Species number

    SizeMytilus

    L cmnumber

    661

    642

    624

    606

    586Size composition of Mytilus galloprovincialis fouling the blocks ( XI, 2001 g)

    565

    5411

    5224

    5025

    4825

    4617

    4413

    4219

    4015

    3825

    3630

    3435

    3267

    30102

    28108

    26120

    24128

    2290

    2079

    1850

    1625Size composition of Mytilus -PZ-MP2001

    1426

    1224

    1021

    840

    646

    443

    218

    1.610

    SizeMytilus

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    L cm

    %

    upperblock

    0

    0.3344481605

    1.0033444816

    0.6688963211

    1.0033444816

    0.6688963211

    1.0033444816

    1.0033444816

    1.6722408027

    1.0033444816

    2.6755852843

    3.0100334448

    4.0133779264

    3.3444816054

    3.6789297659

    4.0133779264

    4.347826087

    3.0100334448

    3.3444816054

    4.6822742475

    5.016722408

    3.3444816054

    3.6789297659

    0.6688963211

    3.6789297659

    2.0066889632

    2.3411371237

    2.0066889632

    3.3444816054

    2.6755852843

    1.3377926421

    2.0066889632

    0.3344481605

    0.3344481605

    0.3344481605

    1.3377926421

    4.347826087

    10.3678929766

    6.3545150502

    0

    mm

    %

    Fouling block - upper side - 14.09.2002- device x 50

    st.1 (US)st.2 (SS)st.3st.4st.5st.6st.7st.8st.9st.10Species compositionAbundanceBiomass

    H(Abund)% frequen

    NSpecies compositionind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageSpecies compositionst.1st.2st.3st.4st.5st.6st.7st.8st.9st.10AverageH(biom)

    1Turbellaria g. sp.00500.0150000005000.14000000001Turbellaria g. sp.0500005000000550.006048441420Turbellaria g. sp.00.0150000.1400000.01550.0000084245

    2Leptoplana sp. St.3,8 - juv000018000.4655500.084000.080013500.5751000.022500.0255501.152Leptoplana sp. St.3 - juv001800550400013501002505505000.038776808570Leptoplana sp. St.3 - juv000.4650.080.0800.5750.020.0251.150.23950.0001059978

    3Prionospio cirrifera44001.0436500.48559001.94518500.2759000.1518500.217500.16529500.8635000.87516500.2653Prionospio cirrifera4400365059001850900185075029503500165027400.1440408182100Prionospio cirrifera1.040.4851.9450.2750.150.210.1650.860.8750.2650.6270.0002552494

    4Heteromastus filiformis1500.31500.031000.261000.0450000500.07500.0253000.495004Heteromastus filiformis- 80%150501001000050503000800.008357485980Heteromastus filiformis0.310.030.260.045000.070.0250.49500.12350.0000576744

    5Nereis succinea - st.3,4- 90%-juv.485038.5251070038.515835040.41555032.62640040.645565050.5840050.05945013.37715015.18520038.1855Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071700.2756262944100Nereis succinea - st.3,4- 90%-juv.38.52538.51540.4132.6240.64550.550.0513.3715.1838.18535.80.0092352074

    6Aricidea claudiae0000500.01500001500.02001000.011000.01006Aricidea claudiae00500015001001000400.004585950550Aricidea claudiae000.015000.0200.010.0100.00550.0000031995

    7Harmothoe reticulata1000.922001.172501.6851000.593504.4753002.835500.395500.42001501.617Harmothoe reticulata - 90% frequency100200250100350300505001501550.014686974590Harmothoe reticulata0.921.171.6850.594.4752.8350.3950.4201.611.410.000531876

    8Harmothoe imbricata500.821000.971501.9452504.092002.315500.7651001.3751000.875501.035500.388Harmothoe imbricata501001502502005010010050501100.010977062100Harmothoe imbricata0.820.971.9454.092.3150.7651.3750.8751.0350.381.4570.0005478438

    9Polydora ciliata-st.4-juv.2000.02513000.04566000.4556500.01517000.24515500.23522000.2059500.1056500.047000.359Polydora ciliata-st.4-juv.2001300660065017001550220095065070016500.099030668100Polydora ciliata-st.4-juv.0.0250.0450.4550.0150.2450.2350.2050.1050.040.350.1720.0000782231

    10Balanus improvisus-mnogo iuv.550103.518250162.5814010013439900121.528100354.173750212192350204.1370800223.682345027.4880150139.27510Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667400.3788560085100Balanus improvisus-mnogo iuv.103.5162.58134121.5354.1212204.13223.6827.48139.275168.22450.0337990013

    11Pisidia longimana- 50 ind. postlarvae2000.7755500.22511501.264500.1851500.0255503.567002.091500.0254001.4953501.5111Pisidia longimana- 50 ind. postlarvae20055011504501505507001504003504650.0365580471100Pisidia longimana- 50 ind. postlarvae0.7750.2251.260.1850.0253.562.090.0251.4951.511.1150.0004302469

    12Upogebia pusilla000000000000500.014500000012Upogebia pusilla0000005000050.000725946710Upogebia pusilla0000000.01450000.001450.0000009148

    13Palaemon elegans00000000000000509.90513Palaemon elegans0000000500050.000725946710Palaemon elegans00000009.905000.99050.0003865298

    14Atanas nitescens-st.4-postlarvae00001001.135500.045501.9051001.16500.6751000.021002.03500.0214Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050600.00652162480Atanas nitescens-st.4-postlarvae001.1350.0451.9051.160.6750.022.030.020.6990.0002817588

    15Alpheus dentipes - larvae501.8700000000000000000015Alpheus dentipes - larvae5000000000050.000725946710Alpheus dentipes - larvae1.870000000000.1870.0000844684

    16Pilumnus hirtellus0000000000100331.92007.355100113.80500509.6716Pilumnus hirtellus00000100200100050450.005081350240Pilumnus hirtellus00000331.97.355113.80509.6746.2730.0114990942

    17Microdeutious gryllotalpa3000.0851000.023500.0800500.040000003000.10017Microdeutious gryllotalpa30010035005000030001100.01097706250Microdeutious gryllotalpa0.0850.020.0800.040000.100.03250.0000167772

    18Melita palmata4000.08965500.4310501.875005000.975501.412001.991500.033500.170018Melita palmata - 80%40055010500500550120015035004750.037195804580Melita palmata0.08960.431.87500.971.41.990.030.1700.695460.0002804621

    19Iphinoe tenella2000.100000000000000000019Iphinoe tenella200000000000200.00249657910Iphinoe tenella0.10000000000.010.0000055968

    20Iphinoe elisae0000500.03000000500.030000500.0320Iphinoe elisae0050000500050150.001935811730Iphinoe elisae000.030000.03000.030.0090.0000050721

    21Corophium bonelli00500.01000000000000000021Corophium bonelli0500000000050.000725946710Corophium bonelli00.01000000000.0010.0000006446

    22Cumella pygmaea euxinica00500.005500.01500000000500.0153500.090022Cumella pygmaea euxinica050500000503500500.005568573730Cumella pygmaea euxinica00.0050.01500000.0150.0900.01250.0000068931

    23Hydrobia acuta5001.06500.16000000500.030000000023Hydrobia acuta50050000500000600.00652162420Hydrobia acuta1.060.160000.0300000.1250.0000583192

    24Rissoa parva0000500.10000000000000024Rissoa parva0050000000050.000725946710Rissoa parva000.100000000.010.0000055968

    25Odostomia pallida005500.4158500.637500.6850000005000.2556500.461000.08525Odostomia pallida05508507500005006501003400.028294054260Odostomia pallida00.4150.630.6850000.2550.460.0850.2530.0001114611

    26Crysalida inserta00005000.161000.02000000500.0151500.0450026Crysalida inserta00500100000501500800.008357485940Crysalida inserta000.160.020000.0150.04500.0240.0000126576

    27Parvicardium exiguum-st.4-juv.003000.92500.2500.16500.105500.3002000.505500.09500.0827Parvicardium exiguum-st.4-juv. - 80%03005050505002005050800.008357485980Parvicardium exiguum-st.4-juv.00.920.20.160.1050.300.5050.090.080.2360.0001045769

    28Cardium edule00000000500.2450000500.19000028Cardium edule000050005000100.001350091420Cardium edule00000.245000.19000.04350.000021988

    29Spisula subtruncata1003800000000000000000029Spisula subtruncata100000000000100.001350091410Spisula subtruncata380000000003.80.0012945226

    30Pitar rudis00000000000000005011.7450030Pitar rudis0000000050050.000725946710Pitar rudis0000000011.74501.17450.000450955

    31Scapharca inaeuqivalvis-juv.001000.47500500.17500000000000031Scapharca inaeuqivalvis-juv.100050000000150.001935811720Scapharca inaeuqivalvis-juv.0.47500.1750000000.0650.0000318932

    32Mytilus galloprovincialis94001770026900372002450040900160502725014600660001885075040170005625013800174501315021960.5156002887032Mytilus galloprovincialis9400269002450016050146001885017000138001315015600169850.4377925648100Mytilus galloprovincialis177003720040900272506600075040562501745021960.52887038862.050.0098227448

    33Molgula euprocta1501.21003002.04500500.08503.72500002000.140033Molgula euprocta150030005050002000750.007906233450Molgula euprocta1.2102.04500.083.725000.1400.720.0002894379

    34Bothrillus schlosseri00013.26500.99000000000006.5903.30534Actinia equina - st.3,4 - juv.00505000055000650.0069886840Bothrillus schlosseri013.2650.99000006.593.3052.4150.0008630655

    35Membranipora denticulata000000000003.5050000000035010Membranipora denticulata000003.50500000.35050.000150203

    36Actinia equina - st.3,4 - juv.0000500.01500.0350000005500.100003630Actinia equina - st.3,4 - juv.000.010.0350000.1000.01450.0000079167

    Total:2160017888.32966350037419.73519235041089.716650027410.525355066405.3810410075652.28522450056519.119510030017814.235120022028.59510470029065.915Total:2160063500192350665005355010410022450010030051200104700982301.600531167Total:17888.329637419.73541089.7127410.5266405.3875652.28556519.119517814.2322028.59529065.91539129.381910.0708464942

    Number of species2 years period (2001 g)Num. Sp. 1 year period(2000 g)Number of species - 1 and 2 years after positioning the blocks

    Blocks%20002001H(A)=1.60

    Vermes925Vermes25Vermes4Vermes49H(B)=0.07

    Crustacea1336.1Crustacea36.1Crustacea1Crustacea113

    Mollusca1027.8Mollusca27.8Mollusca1Mollusca110

    Varia411.1Varia11.1Varia3Varia34

    Total:36100100Total:9Total:936

    speciesnumber% share of various groups (species number)

    Average biomass (10 stations)

    Av. Biomass

    Average abundanceAv. BiomassStructure according to abundance (average 1-10 st)Av. Abundance

    st. 12160017888.3296Vermes39.85

    26350037419.735Vermes12500Crustacea218.25091

    319235041089.71Crustacea68000Mollusca38867.781

    46650027410.52Mollusca17590Varia3.5

    55355066405.38Varia140Total39129.38191

    610410075652.285Total98230

    722450056519.1195

    810030017814.23

    95120022028.595

    1010470029065.915

    Av.9823039129.38191

    Av. Abundance(10 stations)

    ind/m2

    Vermes12500

    Crustacea68000

    Mollusca17590

    Varia140

    Total98230

    Structure according to abundance

    XI, 2000 g.

    Fouling on blocks in 1 year periodAbund.Biomass

    Species compositionind/m2g/m2

    1Leptoplana sp.120.267

    2Harmothoe reticulata521.216

    3Nereis succinea203022.7

    4Polydora ciliata7350.213

    5Balanus improvisus73059.46

    6Mytilus galloprovincialis75306562.5

    7Ctenicella appendiculata70.38

    8Molgula euprocta250.99

    9Bothrillus schlosseri3.12

    Total:111216650.846

    Broj vidove2000 gAv. Abundance

    Vermes42000 g2001 g

    Crustacea1Vermes282912500

    Mollusca1Crustacea73068000

    Varia3Mollusca753017590

    Total:9Varia32140

    Total1112198230

    Abundance per groups in 1 and 2 year periods (1999-2000-2001).

    Average Biomass

    2000 g2001 g

    Vermes24.39639.85

    Crustacea59.46218.25091

    Mollusca6562.538867.781

    Varia4.493.5

    Total6650.84639129.38191

    Biomass per groups in 1 and 2 year periods (1999-2000-2001).

    Dominant species on blocks

    Dominant species on blocks

    B

    Vermes7

    Crustacea5

    Mollusca3

    Species compositionAbundanceVaria1

    Dominant (50-100% frequency)Total16

    st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageN occF%N all stationsAbund 5 stat.D%Species compositionD avgAbundance of dominant species on blocks

    Turbellaria g. sp.05000050000005520PB

    Leptoplana sp. St.3 - juv00180055040001350100250550500701Leptoplana sp. St.3 - juv500Balanus66740Vermes12405

    Prionospio cirrifera4400365059001850900185075029503500165027401002Prionospio cirrifera2740Mytilus16985Crustacea67850

    Heteromastus filiformis- 80%15050100100005050300080703Heteromastus filiformis- 80%80Nereis succinea7170Mollusca17405

    Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071701004Nereis succinea - st.3,4- 90%-juv.7170Prionospio2740Varia75

    Aricidea claudiae0050001500100100040405Harmothoe reticulata - 90% frequency155Polydora ciliata1650Total97735

    Harmothoe reticulata - 90% frequency10020025010035030050500150155906Harmothoe imbricata110Leptoplana500Abundance of dominant species on blocks

    Harmothoe imbricata501001502502005010010050501101007Polydora ciliata-st.4-juv.1650Melita palmata475

    Polydora ciliata-st.4-juv.2001300660065017001550220095065070016501008Balanus improvisus-mnogo iuv.66740Pisidia longimana465

    Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667401009Pisidia longimana- 50 ind. postlarvae465Odostomia pallida340

    Pisidia longimana- 50 ind. postlarvae200550115045015055070015040035046510010Atanas nitescens-st.4-postlarvae - 80%60Harmothoe reticulata155

    Upogebia pusilla0000005000051011Microdeutious gryllotalpa110Microdeutopus gryllotalpa110

    Palaemon elegans0000000500051012Melita palmata - 80%475Harmothoe imbricata110

    Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050608013Odostomia pallida340Parvicardium exiguum80

    Alpheus dentipes - larvae5000000000051014Parvicardium exiguum80Heteromastus filiformis80

    Pilumnus hirtellus00000100200100050454015Mytilus galloprovincialis16985Molgula75

    Microdeutious gryllotalpa30010035005000030001105016Molgula euprocta75Athanas nitescens60

    Melita palmata - 80%400550105005005501200150350047580Total97735

    Iphinoe tenella2000000000002010

    Iphinoe elisae00500005000501530

    Corophium bonelli05000000000510

    Cumella pygmaea euxinica0505000005035005040Abundance of dominant (50-100% frequency) species fouling the blocks

    Hydrobia acuta500500005000006030

    Rissoa parva00500000000510

    Odostomia pallida055085075000050065010034060

    Crysalida inserta005001000005015008040

    Parvicardium exiguum-st.4-juv. - 80%030050505050020050508080

    Cardium edule0000500050001010

    Spisula subtruncata1000000000001010

    Pitar rudis00000000500510

    Scapharca inaeuqivalvis-juv.1000500000001520

    Mytilus galloprovincialis940026900245001605014600188501700013800131501560016985100

    Molgula euprocta1500300050500020007550

    Actinia equina - st.3,4 - juv.005050000550006530

    Total:216006350019235066500535501041002245001003005120010470098230

    0

    0

    0

    0

    0

    0

    0

    0

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0

    0

    0

    0

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    Species number

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    ind / m2

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    g / m2

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Number of species

    Chart6

    2829730753032

    125006800017590140

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / m2

    SizeMytilus

    L cmnumber

    661

    642

    624

    606

    586Size composition of Mytilus galloprovincialis fouling the blocks ( XI, 2001 g)

    565

    5411

    5224

    5025

    4825

    4617

    4413

    4219

    4015

    3825

    3630

    3435

    3267

    30102

    28108

    26120

    24128

    2290

    2079

    1850

    1625Size composition of Mytilus -PZ-MP2001

    1426

    1224

    1021

    840

    646

    443

    218

    1.610

    SizeMytilus

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    L cm

    %

    upperblock

    0

    0.3344481605

    1.0033444816

    0.6688963211

    1.0033444816

    0.6688963211

    1.0033444816

    1.0033444816

    1.6722408027

    1.0033444816

    2.6755852843

    3.0100334448

    4.0133779264

    3.3444816054

    3.6789297659

    4.0133779264

    4.347826087

    3.0100334448

    3.3444816054

    4.6822742475

    5.016722408

    3.3444816054

    3.6789297659

    0.6688963211

    3.6789297659

    2.0066889632

    2.3411371237

    2.0066889632

    3.3444816054

    2.6755852843

    1.3377926421

    2.0066889632

    0.3344481605

    0.3344481605

    0.3344481605

    1.3377926421

    4.347826087

    10.3678929766

    6.3545150502

    0

    mm

    %

    Fouling block - upper side - 14.09.2002- device x 50

    st.1 (US)st.2 (SS)st.3st.4st.5st.6st.7st.8st.9st.10Species compositionAbundanceBiomass

    H(Abund)% frequen

    NSpecies compositionind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageSpecies compositionst.1st.2st.3st.4st.5st.6st.7st.8st.9st.10AverageH(biom)

    1Turbellaria g. sp.00500.0150000005000.14000000001Turbellaria g. sp.0500005000000550.006048441420Turbellaria g. sp.00.0150000.1400000.01550.0000084245

    2Leptoplana sp. St.3,8 - juv000018000.4655500.084000.080013500.5751000.022500.0255501.152Leptoplana sp. St.3 - juv001800550400013501002505505000.038776808570Leptoplana sp. St.3 - juv000.4650.080.0800.5750.020.0251.150.23950.0001059978

    3Prionospio cirrifera44001.0436500.48559001.94518500.2759000.1518500.217500.16529500.8635000.87516500.2653Prionospio cirrifera4400365059001850900185075029503500165027400.1440408182100Prionospio cirrifera1.040.4851.9450.2750.150.210.1650.860.8750.2650.6270.0002552494

    4Heteromastus filiformis1500.31500.031000.261000.0450000500.07500.0253000.495004Heteromastus filiformis- 80%150501001000050503000800.008357485980Heteromastus filiformis0.310.030.260.045000.070.0250.49500.12350.0000576744

    5Nereis succinea - st.3,4- 90%-juv.485038.5251070038.515835040.41555032.62640040.645565050.5840050.05945013.37715015.18520038.1855Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071700.2756262944100Nereis succinea - st.3,4- 90%-juv.38.52538.51540.4132.6240.64550.550.0513.3715.1838.18535.80.0092352074

    6Aricidea claudiae0000500.01500001500.02001000.011000.01006Aricidea claudiae00500015001001000400.004585950550Aricidea claudiae000.015000.0200.010.0100.00550.0000031995

    7Harmothoe reticulata1000.922001.172501.6851000.593504.4753002.835500.395500.42001501.617Harmothoe reticulata - 90% frequency100200250100350300505001501550.014686974590Harmothoe reticulata0.921.171.6850.594.4752.8350.3950.4201.611.410.000531876

    8Harmothoe imbricata500.821000.971501.9452504.092002.315500.7651001.3751000.875501.035500.388Harmothoe imbricata501001502502005010010050501100.010977062100Harmothoe imbricata0.820.971.9454.092.3150.7651.3750.8751.0350.381.4570.0005478438

    9Polydora ciliata-st.4-juv.2000.02513000.04566000.4556500.01517000.24515500.23522000.2059500.1056500.047000.359Polydora ciliata-st.4-juv.2001300660065017001550220095065070016500.099030668100Polydora ciliata-st.4-juv.0.0250.0450.4550.0150.2450.2350.2050.1050.040.350.1720.0000782231

    10Balanus improvisus-mnogo iuv.550103.518250162.5814010013439900121.528100354.173750212192350204.1370800223.682345027.4880150139.27510Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667400.3788560085100Balanus improvisus-mnogo iuv.103.5162.58134121.5354.1212204.13223.6827.48139.275168.22450.0337990013

    11Pisidia longimana- 50 ind. postlarvae2000.7755500.22511501.264500.1851500.0255503.567002.091500.0254001.4953501.5111Pisidia longimana- 50 ind. postlarvae20055011504501505507001504003504650.0365580471100Pisidia longimana- 50 ind. postlarvae0.7750.2251.260.1850.0253.562.090.0251.4951.511.1150.0004302469

    12Upogebia pusilla000000000000500.014500000012Upogebia pusilla0000005000050.000725946710Upogebia pusilla0000000.01450000.001450.0000009148

    13Palaemon elegans00000000000000509.90513Palaemon elegans0000000500050.000725946710Palaemon elegans00000009.905000.99050.0003865298

    14Atanas nitescens-st.4-postlarvae00001001.135500.045501.9051001.16500.6751000.021002.03500.0214Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050600.00652162480Atanas nitescens-st.4-postlarvae001.1350.0451.9051.160.6750.022.030.020.6990.0002817588

    15Alpheus dentipes - larvae501.8700000000000000000015Alpheus dentipes - larvae5000000000050.000725946710Alpheus dentipes - larvae1.870000000000.1870.0000844684

    16Pilumnus hirtellus0000000000100331.92007.355100113.80500509.6716Pilumnus hirtellus00000100200100050450.005081350240Pilumnus hirtellus00000331.97.355113.80509.6746.2730.0114990942

    17Microdeutious gryllotalpa3000.0851000.023500.0800500.040000003000.10017Microdeutious gryllotalpa30010035005000030001100.01097706250Microdeutious gryllotalpa0.0850.020.0800.040000.100.03250.0000167772

    18Melita palmata4000.08965500.4310501.875005000.975501.412001.991500.033500.170018Melita palmata - 80%40055010500500550120015035004750.037195804580Melita palmata0.08960.431.87500.971.41.990.030.1700.695460.0002804621

    19Iphinoe tenella2000.100000000000000000019Iphinoe tenella200000000000200.00249657910Iphinoe tenella0.10000000000.010.0000055968

    20Iphinoe elisae0000500.03000000500.030000500.0320Iphinoe elisae0050000500050150.001935811730Iphinoe elisae000.030000.03000.030.0090.0000050721

    21Corophium bonelli00500.01000000000000000021Corophium bonelli0500000000050.000725946710Corophium bonelli00.01000000000.0010.0000006446

    22Cumella pygmaea euxinica00500.005500.01500000000500.0153500.090022Cumella pygmaea euxinica050500000503500500.005568573730Cumella pygmaea euxinica00.0050.01500000.0150.0900.01250.0000068931

    23Hydrobia acuta5001.06500.16000000500.030000000023Hydrobia acuta50050000500000600.00652162420Hydrobia acuta1.060.160000.0300000.1250.0000583192

    24Rissoa parva0000500.10000000000000024Rissoa parva0050000000050.000725946710Rissoa parva000.100000000.010.0000055968

    25Odostomia pallida005500.4158500.637500.6850000005000.2556500.461000.08525Odostomia pallida05508507500005006501003400.028294054260Odostomia pallida00.4150.630.6850000.2550.460.0850.2530.0001114611

    26Crysalida inserta00005000.161000.02000000500.0151500.0450026Crysalida inserta00500100000501500800.008357485940Crysalida inserta000.160.020000.0150.04500.0240.0000126576

    27Parvicardium exiguum-st.4-juv.003000.92500.2500.16500.105500.3002000.505500.09500.0827Parvicardium exiguum-st.4-juv. - 80%03005050505002005050800.008357485980Parvicardium exiguum-st.4-juv.00.920.20.160.1050.300.5050.090.080.2360.0001045769

    28Cardium edule00000000500.2450000500.19000028Cardium edule000050005000100.001350091420Cardium edule00000.245000.19000.04350.000021988

    29Spisula subtruncata1003800000000000000000029Spisula subtruncata100000000000100.001350091410Spisula subtruncata380000000003.80.0012945226

    30Pitar rudis00000000000000005011.7450030Pitar rudis0000000050050.000725946710Pitar rudis0000000011.74501.17450.000450955

    31Scapharca inaeuqivalvis-juv.001000.47500500.17500000000000031Scapharca inaeuqivalvis-juv.100050000000150.001935811720Scapharca inaeuqivalvis-juv.0.47500.1750000000.0650.0000318932

    32Mytilus galloprovincialis94001770026900372002450040900160502725014600660001885075040170005625013800174501315021960.5156002887032Mytilus galloprovincialis9400269002450016050146001885017000138001315015600169850.4377925648100Mytilus galloprovincialis177003720040900272506600075040562501745021960.52887038862.050.0098227448

    33Molgula euprocta1501.21003002.04500500.08503.72500002000.140033Molgula euprocta150030005050002000750.007906233450Molgula euprocta1.2102.04500.083.725000.1400.720.0002894379

    34Bothrillus schlosseri00013.26500.99000000000006.5903.30534Actinia equina - st.3,4 - juv.00505000055000650.0069886840Bothrillus schlosseri013.2650.99000006.593.3052.4150.0008630655

    35Membranipora denticulata000000000003.5050000000035010Membranipora denticulata000003.50500000.35050.000150203

    36Actinia equina - st.3,4 - juv.0000500.01500.0350000005500.100003630Actinia equina - st.3,4 - juv.000.010.0350000.1000.01450.0000079167

    Total:2160017888.32966350037419.73519235041089.716650027410.525355066405.3810410075652.28522450056519.119510030017814.235120022028.59510470029065.915Total:2160063500192350665005355010410022450010030051200104700982301.600531167Total:17888.329637419.73541089.7127410.5266405.3875652.28556519.119517814.2322028.59529065.91539129.381910.0708464942

    Number of species2 years period (2001 g)Num. Sp. 1 year period(2000 g)Number of species - 1 and 2 years after positioning the blocks

    Blocks%20002001H(A)=1.60

    Vermes925Vermes25Vermes4Vermes49H(B)=0.07

    Crustacea1336.1Crustacea36.1Crustacea1Crustacea113

    Mollusca1027.8Mollusca27.8Mollusca1Mollusca110

    Varia411.1Varia11.1Varia3Varia34

    Total:36100100Total:9Total:936

    speciesnumber% share of various groups (species number)

    Average biomass (10 stations)

    Av. Biomass

    Average abundanceAv. BiomassStructure according to abundance (average 1-10 st)Av. Abundance

    st. 12160017888.3296Vermes39.85

    26350037419.735Vermes12500Crustacea218.25091

    319235041089.71Crustacea68000Mollusca38867.781

    46650027410.52Mollusca17590Varia3.5

    55355066405.38Varia140Total39129.38191

    610410075652.285Total98230

    722450056519.1195

    810030017814.23

    95120022028.595

    1010470029065.915

    Av.9823039129.38191

    Av. Abundance(10 stations)

    ind/m2

    Vermes12500

    Crustacea68000

    Mollusca17590

    Varia140

    Total98230

    Structure according to abundance

    XI, 2000 g.

    Fouling on blocks in 1 year periodAbund.Biomass

    Species compositionind/m2g/m2

    1Leptoplana sp.120.267

    2Harmothoe reticulata521.216

    3Nereis succinea203022.7

    4Polydora ciliata7350.213

    5Balanus improvisus73059.46

    6Mytilus galloprovincialis75306562.5

    7Ctenicella appendiculata70.38

    8Molgula euprocta250.99

    9Bothrillus schlosseri3.12

    Total:111216650.846

    Broj vidove2000 gAv. Abundance

    Vermes420002001

    Crustacea1Vermes282912500

    Mollusca1Crustacea73068000

    Varia3Mollusca753017590

    Total:9Varia32140

    Total1112198230

    Abundance per groups in 1 and 2 year periods (1999-2000-2001).

    Average Biomass

    20002001

    Vermes24.39639.85

    Crustacea59.46218.25091

    Mollusca6562.538867.781

    Varia4.493.5

    Total6650.84639129.38191

    Biomass per groups in 1 and 2 year periods (1999-2000-2001).

    Dominant species on blocks

    Dominant species on blocks

    B

    Vermes7

    Crustacea5

    Mollusca3

    Species compositionAbundanceVaria1

    Dominant (50-100% frequency)Total16

    st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageN occF%N all stationsAbund 5 stat.D%Species compositionD avgAbundance of dominant species on blocks

    Turbellaria g. sp.05000050000005520PB

    Leptoplana sp. St.3 - juv00180055040001350100250550500701Leptoplana sp. St.3 - juv500B. improvisus66740Vermes12405

    Prionospio cirrifera4400365059001850900185075029503500165027401002Prionospio cirrifera2740M. galloprovincialis16985Crustacea67850

    Heteromastus filiformis- 80%15050100100005050300080703Heteromastus filiformis- 80%80N. succinea7170Mollusca17405

    Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071701004Nereis succinea - st.3,4- 90%-juv.7170Pr.cirrifera2740Varia75

    Aricidea claudiae0050001500100100040405Harmothoe reticulata - 90% frequency155P. ciliata1650Total97735

    Harmothoe reticulata - 90% frequency10020025010035030050500150155906Harmothoe imbricata110Leptoplana g. sp.500Abundance of dominant species on blocks

    Harmothoe imbricata501001502502005010010050501101007Polydora ciliata-st.4-juv.1650M. palmata475

    Polydora ciliata-st.4-juv.2001300660065017001550220095065070016501008Balanus improvisus-mnogo iuv.66740P. longimana465

    Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667401009Pisidia longimana- 50 ind. postlarvae465Odostomia pallida340

    Pisidia longimana- 50 ind. postlarvae200550115045015055070015040035046510010Atanas nitescens-st.4-postlarvae - 80%60Harmothoe reticulata155

    Upogebia pusilla0000005000051011Microdeutious gryllotalpa110Microdeutopus gryllotalpa110

    Palaemon elegans0000000500051012Melita palmata - 80%475Harmothoe imbricata110

    Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050608013Odostomia pallida340Parvicardium exiguum80

    Alpheus dentipes - larvae5000000000051014Parvicardium exiguum80Heteromastus filiformis80

    Pilumnus hirtellus00000100200100050454015Mytilus galloprovincialis16985Molgula75

    Microdeutious gryllotalpa30010035005000030001105016Molgula euprocta75Athanas nitescens60

    Melita palmata - 80%400550105005005501200150350047580Total97735

    Iphinoe tenella2000000000002010

    Iphinoe elisae00500005000501530

    Corophium bonelli05000000000510

    Cumella pygmaea euxinica0505000005035005040Abundance of dominant (50-100% frequency) species fouling the blocks

    Hydrobia acuta500500005000006030

    Rissoa parva00500000000510

    Odostomia pallida055085075000050065010034060

    Crysalida inserta005001000005015008040

    Parvicardium exiguum-st.4-juv. - 80%030050505050020050508080

    Cardium edule0000500050001010

    Spisula subtruncata1000000000001010

    Pitar rudis00000000500510

    Scapharca inaeuqivalvis-juv.1000500000001520

    Mytilus galloprovincialis940026900245001605014600188501700013800131501560016985100

    Molgula euprocta1500300050500020007550

    Actinia equina - st.3,4 - juv.005050000550006530

    Total:216006350019235066500535501041002245001003005120010470098230

    0

    0

    0

    0

    0

    0

    0

    0

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0

    0

    0

    0

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    Species number

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / m2

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Number of species

    0

    0

    0

    0

    0

    0

    0

    0

    ind/sq.m

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / sq.m

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / sq.m

    Chart8

    24.39659.466562.54.49

    39.85218.2509138867.7813.5

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / sq.m

    SizeMytilus

    L cmnumber

    661

    642

    624

    606

    586Size composition of Mytilus galloprovincialis fouling the blocks ( XI, 2001 g)

    565

    5411

    5224

    5025

    4825

    4617

    4413

    4219

    4015

    3825

    3630

    3435

    3267

    30102

    28108

    26120

    24128

    2290

    2079

    1850

    1625Size composition of Mytilus -PZ-MP2001

    1426

    1224

    1021

    840

    646

    443

    218

    1.610

    SizeMytilus

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    L cm

    %

    upperblock

    0

    0.3344481605

    1.0033444816

    0.6688963211

    1.0033444816

    0.6688963211

    1.0033444816

    1.0033444816

    1.6722408027

    1.0033444816

    2.6755852843

    3.0100334448

    4.0133779264

    3.3444816054

    3.6789297659

    4.0133779264

    4.347826087

    3.0100334448

    3.3444816054

    4.6822742475

    5.016722408

    3.3444816054

    3.6789297659

    0.6688963211

    3.6789297659

    2.0066889632

    2.3411371237

    2.0066889632

    3.3444816054

    2.6755852843

    1.3377926421

    2.0066889632

    0.3344481605

    0.3344481605

    0.3344481605

    1.3377926421

    4.347826087

    10.3678929766

    6.3545150502

    0

    mm

    %

    Fouling block - upper side - 14.09.2002- device x 50

    st.1 (US)st.2 (SS)st.3st.4st.5st.6st.7st.8st.9st.10Species compositionAbundanceBiomass

    H(Abund)% frequen

    NSpecies compositionind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2ind/m2g/m2st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageSpecies compositionst.1st.2st.3st.4st.5st.6st.7st.8st.9st.10AverageH(biom)

    1Turbellaria g. sp.00500.0150000005000.14000000001Turbellaria g. sp.0500005000000550.006048441420Turbellaria g. sp.00.0150000.1400000.01550.0000084245

    2Leptoplana sp. St.3,8 - juv000018000.4655500.084000.080013500.5751000.022500.0255501.152Leptoplana sp. St.3 - juv001800550400013501002505505000.038776808570Leptoplana sp. St.3 - juv000.4650.080.0800.5750.020.0251.150.23950.0001059978

    3Prionospio cirrifera44001.0436500.48559001.94518500.2759000.1518500.217500.16529500.8635000.87516500.2653Prionospio cirrifera4400365059001850900185075029503500165027400.1440408182100Prionospio cirrifera1.040.4851.9450.2750.150.210.1650.860.8750.2650.6270.0002552494

    4Heteromastus filiformis1500.31500.031000.261000.0450000500.07500.0253000.495004Heteromastus filiformis- 80%150501001000050503000800.008357485980Heteromastus filiformis0.310.030.260.045000.070.0250.49500.12350.0000576744

    5Nereis succinea - st.3,4- 90%-juv.485038.5251070038.515835040.41555032.62640040.645565050.5840050.05945013.37715015.18520038.1855Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071700.2756262944100Nereis succinea - st.3,4- 90%-juv.38.52538.51540.4132.6240.64550.550.0513.3715.1838.18535.80.0092352074

    6Aricidea claudiae0000500.01500001500.02001000.011000.01006Aricidea claudiae00500015001001000400.004585950550Aricidea claudiae000.015000.0200.010.0100.00550.0000031995

    7Harmothoe reticulata1000.922001.172501.6851000.593504.4753002.835500.395500.42001501.617Harmothoe reticulata - 90% frequency100200250100350300505001501550.014686974590Harmothoe reticulata0.921.171.6850.594.4752.8350.3950.4201.611.410.000531876

    8Harmothoe imbricata500.821000.971501.9452504.092002.315500.7651001.3751000.875501.035500.388Harmothoe imbricata501001502502005010010050501100.010977062100Harmothoe imbricata0.820.971.9454.092.3150.7651.3750.8751.0350.381.4570.0005478438

    9Polydora ciliata-st.4-juv.2000.02513000.04566000.4556500.01517000.24515500.23522000.2059500.1056500.047000.359Polydora ciliata-st.4-juv.2001300660065017001550220095065070016500.099030668100Polydora ciliata-st.4-juv.0.0250.0450.4550.0150.2450.2350.2050.1050.040.350.1720.0000782231

    10Balanus improvisus-mnogo iuv.550103.518250162.5814010013439900121.528100354.173750212192350204.1370800223.682345027.4880150139.27510Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667400.3788560085100Balanus improvisus-mnogo iuv.103.5162.58134121.5354.1212204.13223.6827.48139.275168.22450.0337990013

    11Pisidia longimana- 50 ind. postlarvae2000.7755500.22511501.264500.1851500.0255503.567002.091500.0254001.4953501.5111Pisidia longimana- 50 ind. postlarvae20055011504501505507001504003504650.0365580471100Pisidia longimana- 50 ind. postlarvae0.7750.2251.260.1850.0253.562.090.0251.4951.511.1150.0004302469

    12Upogebia pusilla000000000000500.014500000012Upogebia pusilla0000005000050.000725946710Upogebia pusilla0000000.01450000.001450.0000009148

    13Palaemon elegans00000000000000509.90513Palaemon elegans0000000500050.000725946710Palaemon elegans00000009.905000.99050.0003865298

    14Atanas nitescens-st.4-postlarvae00001001.135500.045501.9051001.16500.6751000.021002.03500.0214Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050600.00652162480Atanas nitescens-st.4-postlarvae001.1350.0451.9051.160.6750.022.030.020.6990.0002817588

    15Alpheus dentipes - larvae501.8700000000000000000015Alpheus dentipes - larvae5000000000050.000725946710Alpheus dentipes - larvae1.870000000000.1870.0000844684

    16Pilumnus hirtellus0000000000100331.92007.355100113.80500509.6716Pilumnus hirtellus00000100200100050450.005081350240Pilumnus hirtellus00000331.97.355113.80509.6746.2730.0114990942

    17Microdeutious gryllotalpa3000.0851000.023500.0800500.040000003000.10017Microdeutious gryllotalpa30010035005000030001100.01097706250Microdeutious gryllotalpa0.0850.020.0800.040000.100.03250.0000167772

    18Melita palmata4000.08965500.4310501.875005000.975501.412001.991500.033500.170018Melita palmata - 80%40055010500500550120015035004750.037195804580Melita palmata0.08960.431.87500.971.41.990.030.1700.695460.0002804621

    19Iphinoe tenella2000.100000000000000000019Iphinoe tenella200000000000200.00249657910Iphinoe tenella0.10000000000.010.0000055968

    20Iphinoe elisae0000500.03000000500.030000500.0320Iphinoe elisae0050000500050150.001935811730Iphinoe elisae000.030000.03000.030.0090.0000050721

    21Corophium bonelli00500.01000000000000000021Corophium bonelli0500000000050.000725946710Corophium bonelli00.01000000000.0010.0000006446

    22Cumella pygmaea euxinica00500.005500.01500000000500.0153500.090022Cumella pygmaea euxinica050500000503500500.005568573730Cumella pygmaea euxinica00.0050.01500000.0150.0900.01250.0000068931

    23Hydrobia acuta5001.06500.16000000500.030000000023Hydrobia acuta50050000500000600.00652162420Hydrobia acuta1.060.160000.0300000.1250.0000583192

    24Rissoa parva0000500.10000000000000024Rissoa parva0050000000050.000725946710Rissoa parva000.100000000.010.0000055968

    25Odostomia pallida005500.4158500.637500.6850000005000.2556500.461000.08525Odostomia pallida05508507500005006501003400.028294054260Odostomia pallida00.4150.630.6850000.2550.460.0850.2530.0001114611

    26Crysalida inserta00005000.161000.02000000500.0151500.0450026Crysalida inserta00500100000501500800.008357485940Crysalida inserta000.160.020000.0150.04500.0240.0000126576

    27Parvicardium exiguum-st.4-juv.003000.92500.2500.16500.105500.3002000.505500.09500.0827Parvicardium exiguum-st.4-juv. - 80%03005050505002005050800.008357485980Parvicardium exiguum-st.4-juv.00.920.20.160.1050.300.5050.090.080.2360.0001045769

    28Cardium edule00000000500.2450000500.19000028Cardium edule000050005000100.001350091420Cardium edule00000.245000.19000.04350.000021988

    29Spisula subtruncata1003800000000000000000029Spisula subtruncata100000000000100.001350091410Spisula subtruncata380000000003.80.0012945226

    30Pitar rudis00000000000000005011.7450030Pitar rudis0000000050050.000725946710Pitar rudis0000000011.74501.17450.000450955

    31Scapharca inaeuqivalvis-juv.001000.47500500.17500000000000031Scapharca inaeuqivalvis-juv.100050000000150.001935811720Scapharca inaeuqivalvis-juv.0.47500.1750000000.0650.0000318932

    32Mytilus galloprovincialis94001770026900372002450040900160502725014600660001885075040170005625013800174501315021960.5156002887032Mytilus galloprovincialis9400269002450016050146001885017000138001315015600169850.4377925648100Mytilus galloprovincialis177003720040900272506600075040562501745021960.52887038862.050.0098227448

    33Molgula euprocta1501.21003002.04500500.08503.72500002000.140033Molgula euprocta150030005050002000750.007906233450Molgula euprocta1.2102.04500.083.725000.1400.720.0002894379

    34Bothrillus schlosseri00013.26500.99000000000006.5903.30534Actinia equina - st.3,4 - juv.00505000055000650.0069886840Bothrillus schlosseri013.2650.99000006.593.3052.4150.0008630655

    35Membranipora denticulata000000000003.5050000000035010Membranipora denticulata000003.50500000.35050.000150203

    36Actinia equina - st.3,4 - juv.0000500.01500.0350000005500.100003630Actinia equina - st.3,4 - juv.000.010.0350000.1000.01450.0000079167

    Total:2160017888.32966350037419.73519235041089.716650027410.525355066405.3810410075652.28522450056519.119510030017814.235120022028.59510470029065.915Total:2160063500192350665005355010410022450010030051200104700982301.600531167Total:17888.329637419.73541089.7127410.5266405.3875652.28556519.119517814.2322028.59529065.91539129.381910.0708464942

    Number of species2 years period (2001 g)Num. Sp. 1 year period(2000 g)Number of species - 1 and 2 years after positioning the blocks

    Blocks%20002001H(A)=1.60

    Vermes925Vermes25Vermes4Vermes49H(B)=0.07

    Crustacea1336.1Crustacea36.1Crustacea1Crustacea113

    Mollusca1027.8Mollusca27.8Mollusca1Mollusca110

    Varia411.1Varia11.1Varia3Varia34

    Total:36100100Total:9Total:936

    speciesnumber% share of various groups (species number)

    Average biomass (10 stations)

    Av. Biomass

    Average abundanceAv. BiomassStructure according to abundance (average 1-10 st)Av. Abundance

    st. 12160017888.3296Vermes39.85

    26350037419.735Vermes12500Crustacea218.25091

    319235041089.71Crustacea68000Mollusca38867.781

    46650027410.52Mollusca17590Varia3.5

    55355066405.38Varia140Total39129.38191

    610410075652.285Total98230

    722450056519.1195

    810030017814.23

    95120022028.595

    1010470029065.915

    Av.9823039129.38191

    Av. Abundance(10 stations)

    ind/m2

    Vermes12500

    Crustacea68000

    Mollusca17590

    Varia140

    Total98230

    Structure according to abundance

    XI, 2000 g.

    Fouling on blocks in 1 year periodAbund.Biomass

    Species compositionind/m2g/m2

    1Leptoplana sp.120.267

    2Harmothoe reticulata521.216

    3Nereis succinea203022.7

    4Polydora ciliata7350.213

    5Balanus improvisus73059.46

    6Mytilus galloprovincialis75306562.5

    7Ctenicella appendiculata70.38

    8Molgula euprocta250.99

    9Bothrillus schlosseri3.12

    Total:111216650.846

    Broj vidove2000 gAv. Abundance

    Vermes420002001

    Crustacea1Vermes282912500

    Mollusca1Crustacea73068000

    Varia3Mollusca753017590

    Total:9Varia32140

    Total1112198230

    Abundance per groups in 1 and 2 year periods (1999-2000-2001).

    Average Biomass

    20002001

    Vermes24.39639.85

    Crustacea59.46218.25091

    Mollusca6562.538867.781

    Varia4.493.5

    Total6650.84639129.38191

    Biomass per groups in 1 and 2 year periods (1999-2000-2001).

    Dominant species on blocks

    Dominant species on blocks

    B

    Vermes7

    Crustacea5

    Mollusca3

    Species compositionAbundanceVaria1

    Dominant (50-100% frequency)Total16

    st.1st.2st.3st.4st.5st.6st.7st.8st.9st.10averageN occF%N all stationsAbund 5 stat.D%Species compositionD avgAbundance of dominant species on blocks

    Turbellaria g. sp.05000050000005520PB

    Leptoplana sp. St.3 - juv00180055040001350100250550500701Leptoplana sp. St.3 - juv500B. improvisus66740Vermes12405

    Prionospio cirrifera4400365059001850900185075029503500165027401002Prionospio cirrifera2740M. galloprovincialis16985Crustacea67850

    Heteromastus filiformis- 80%15050100100005050300080703Heteromastus filiformis- 80%80N. succinea7170Mollusca17405

    Nereis succinea - st.3,4- 90%-juv.4850107008350555064005650840094507150520071701004Nereis succinea - st.3,4- 90%-juv.7170Pr.cirrifera2740Varia75

    Aricidea claudiae0050001500100100040405Harmothoe reticulata - 90% frequency155P. ciliata1650Total97735

    Harmothoe reticulata - 90% frequency10020025010035030050500150155906Harmothoe imbricata110Leptoplana g. sp.500Abundance of dominant species on blocks

    Harmothoe imbricata501001502502005010010050501101007Polydora ciliata-st.4-juv.1650M. palmata475

    Polydora ciliata-st.4-juv.2001300660065017001550220095065070016501008Balanus improvisus-mnogo iuv.66740P. longimana465

    Balanus improvisus-mnogo iuv.55018250140100399002810073750192350708002345080150667401009Pisidia longimana- 50 ind. postlarvae465Odostomia pallida340

    Pisidia longimana- 50 ind. postlarvae200550115045015055070015040035046510010Atanas nitescens-st.4-postlarvae - 80%60Harmothoe reticulata155

    Upogebia pusilla0000005000051011Microdeutious gryllotalpa110Microdeutopus gryllotalpa110

    Palaemon elegans0000000500051012Melita palmata - 80%475Harmothoe imbricata110

    Atanas nitescens-st.4-postlarvae - 80%0010050501005010010050608013Odostomia pallida340Parvicardium exiguum80

    Alpheus dentipes - larvae5000000000051014Parvicardium exiguum80Heteromastus filiformis80

    Pilumnus hirtellus00000100200100050454015Mytilus galloprovincialis16985Molgula75

    Microdeutious gryllotalpa30010035005000030001105016Molgula euprocta75Athanas nitescens60

    Melita palmata - 80%400550105005005501200150350047580Total97735

    Iphinoe tenella2000000000002010

    Iphinoe elisae00500005000501530

    Corophium bonelli05000000000510

    Cumella pygmaea euxinica0505000005035005040Abundance of dominant (50-100% frequency) species fouling the blocks

    Hydrobia acuta500500005000006030

    Rissoa parva00500000000510

    Odostomia pallida055085075000050065010034060

    Crysalida inserta005001000005015008040

    Parvicardium exiguum-st.4-juv. - 80%030050505050020050508080

    Cardium edule0000500050001010

    Spisula subtruncata1000000000001010

    Pitar rudis00000000500510

    Scapharca inaeuqivalvis-juv.1000500000001520

    Mytilus galloprovincialis940026900245001605014600188501700013800131501560016985100

    Molgula euprocta1500300050500020007550

    Actinia equina - st.3,4 - juv.005050000550006530

    Total:216006350019235066500535501041002245001003005120010470098230

    0

    0

    0

    0

    0

    0

    0

    0

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    0

    0

    0

    0

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    Species number

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / m2

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    ind / m2

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Number of species

    0

    0

    0

    0

    0

    0

    0

    0

    ind/sq.m

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / sq.m

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Varia

    Years

    ind / sq.m

  • M. galloprovincialis . . 1984 . .

  • (Konsulov, 1980-Pat. N 28585). 1/ 2/ 3/ (, )4/ Rapana 5/ - 6/ . ( ) ( )

  • .

  • 1993 . . 50 ( ) .

  • , .

  • 2003

  • . ( C ) (R ) (U) ( S ) ( M ) ( O )

  • . : ( C ), ( U ), ( O ), ( R ), ( M ) ( S )-o R (58), M (47), S (43). , (25 24 ), - (20 ).

    Chart3

    101013

    9115U

    1154O

    242464

    1811171

    201391

    Vermes

    Crustacea

    Mollusca

    Miscellaneous

    Species number

    Sheet2

    CUORMS

    Vermes10911241820

    Crustacea10115241113

    Mollusca1546179

    Miscellaneous3411

    242520584743

    Pisces larvae2574

    Sheet2

    0000

    0000

    0000

    0000

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Miscellaneous

    Species number

    Dalboka1

    CCCCCUUUUUOOOOORRRRRMMMMMSSSS

    Leptoplana sp.155312191386835163320080602000000004950000003000000

    Turbellaria g. sp.00000000000000060012000108002100000000000

    Nemertini g. sp.30063791340225003764000000001000170045080014001000005003000100200

    Nematoda g. sp.000000000000800000000000000000

    Aricidea claudiae000000000000000000007050402904400000

    Capitella capitata0000004000100000036000000000000000

    Capitomastus minimus000002000300600400200245015001500020000045020037001600

    Eteone picta000000000000000000015000000100000

    Exogone gemmifera000000000000000000050000000000

    Glycera alba000000000000000000000000050000

    Grubea clavata0000000000000003600001000000000000

    Grubea limbata0000000000000000350450000000050100150200

    Harmothoe imbricata000000000000000501005010025000001050000

    Harmothoe reticulata1172171841261192000102000000000000010000000

    Heteromastus filiformis000000000002020125000050008509208301601500360450200160380

    Leiochone clipeata000000000000000000002000000000

    Lisidice nineta000000000000000000050000000000

    Melinna pal0000000000000200000000810270428081205200000

    Microphthalmus shcelcovi00000000000000000000000000000

    Microphthalmus fragilis0000000000000000000000000000800

    Nephthys hombergii0000000000200000000008080905801600000

    Nereis zonata13226105711189510417978500000000002450700094507500000050000

    Nereis succinea551013451628424080120601602020060020014004200505047001850080203010200300200160

    Nerine cirratulus000000000000000005000000000000

    Oridia armandi000000000000000000006005010100000

    Pectinaria coreni0000000000000000000010002000000

    Perinereis cultrifera00000000000000016000000000000000

    Platynereis dumerilii468276768557256412835127504030506000000805054502550167007000000015000100

    Phyllodoce mucosa0000000000000000000030020000000

    Phyllodoce tuberculata14537851119916126340000000000250020030050000000000

    Polycirrus jubatus000000000000000505000000000300000

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    Polygordius neapolitanus00000000000000000000000002006018020

    Prionospio cirrifera117001109860140208011080802000502000007501300001102750100200860

    Pygospio elegans0000000000000000200005020080010001000

    Spio filicornis00000000000040120800500050004010000360018005700400

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    Syllides longicirrata0000000000000000000000000250000

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    Syllis prolifera0000000000000003500000000000000

    Oligochaeta g.sp.000001004020604013601020280610013007300250004005900390520309209100300680310

    Athanas nitescens2171171671872340000000000000500000000000

    Ampelisca di0000000000000050000015040101020708050900100

    Amphithoe vaillanti21242467612959118658975000000000050350010500000000000

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    Apherusa bispinosa000000000000000200150000000000000

    Balanus improvisus3792688516733128592178034014808601100164020012000260091003200201001865004501020204001600190060

    Brahinotus sexdentatus000002001010200000000000000000000

    Caprella acantifera0000000000000001000503000000000000

    Coremapus versiculatus000000000000000000000000100000

    Corophium bonelli45102658353891006090110120000000506450150503000000000

    Corophium runcicorne0000000000000000000000500400000

    Cumella limicola00000000000000011001501500100000000000

    Cumella pygmaea euxinica00000000000000020050010050000000000

    Dexamine0000000000000000050500000000000

    Diogenes pugilator0000080602040100000000000000000506040120

    Idotea baltica00000060030200000005000000000001000

    Iphinoe elisae00000000000000300000045040080004502010060

    Iphinoe tenella00000000000000000000000000000

    Jassa ocia217010517614700000000007009506009001100000000000

    Leptochelia savignyi0000000000000001100500650100000000000

    Melita palmata3692489412621044587056321340100220018027000008001500245039501100000001800000

    Marinogammarus olivii011764863868020401000000000000000000000

    Microdeutopus gryllotalpa31813238312804029245301947201560122011017400000080014009350149002300000002400000

    Microdeutopuis damnoniensis000000000000000150020000000000000

    Naesa bidentata00000000000000017001005000000000000

    Nototropis guttatus000000000000000000000000050180100100

    Orhestia bottae0000020202010200000000000000000000

    Orhestia montague0000000200100000000000000000000

    Perioculodes longimanus00000000000000000000003050100000

    Pilumnus hirtellus451334392264413000000000010050501001500000050204010

    Pisidia longimana451451423417563000000000050550550650360000001001002060

    Phthysica marina0000000000000000000000017000000

    Pseudocuma longicornis0000000000000000500003010000100200300100

    Pseudocuma graciloides0000000000000000000000000150500100

    Rhithropanopeus harrizi000000000000000500000000000000

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    Stenothoe monoculoides0000000000000002600460051006200380000000000200

    Tanais cavolini000000000000000005000000000000

    Upogebia pusilla000002000100020000000050000000000

    Xantho poressa0000000000000000010000000000000

    Abra alba00000000002000000000000010100000

    Abra nitida000000000000000000000002000000

    Abra renieri0000000000000000000060200000000

    Calyptraea chinensis00000000000000000000000000000

    Cardium edule0000020203004000000000003090013010502010010

    Chamelea gallina000000000000000000000000080033001100640

    Cylichnina variabilis00000000000000000000200800400100500

    Hydrobia acuta0000000000000000000500259000003001000

    Lentidium mediterraneum0000000000000000000000000960400186006800

    Moerella tenuis00000000000000000000010000010000

    Mya arenaria0000000000000000000000102000000

    Mytilus galloprovincialis1801763636262709212201472143568000200600110200000310022002700230028001008010702000000

    Mytilaster lineatus000002400901201600000065001700275010750134000000010000100

    Nana donovani0000000000000000000250000000000

    Nassa reticulata0000024020018011026004000000003501010000500000

    Odostomia pallida0000000000000000050000001000000

    Parvicardium exiguum000000000000000000010000105000000

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    Pitar rudis000000000000000000000001000000

    Rapana thomasiana000000402010300000000000000000000

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    Scapharca inaequivalvis0000000000240000000000020020400000

    Spisula subtruncata00000000000000000000300270250200000

    Sphenia sp.0000000000000000000001000000000

    Actinia equina00000000000000000100050000000000

    Ctenicella apendiculata0000000000000000000100000000000

    Molguls euprocta00000000000000000000000000000

    Chironomidae larvae0000000000000006000000000000000

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    Phoronis euxinicola0000000000000000000034002102600770010000

  • Vermes . (11.6%), Crustacea (70.1%).70.196.0

    Chart6

    75329118325249845.20

    44626927040

    5330156640

    32380274829800484

    58062441188784

    91922464685820

    Vermes

    Crustacea

    Mollusca

    Miscellaneous

    Sheet2

    CUORMS

    Vermes10911241820

    Crustacea10115241113

    Mollusca1546179

    Miscellaneous3411

    242520584743

    Pisces larvae2574

    Sheet2

    0000

    0000

    0000

    0000

    0000

    0000

    Vermes

    Crustacea

    Mollusca

    Miscellaneous

    Species number

    Dalboka1

    CCCCCUUUUUOOOOORRRRRMMMMMSSSS

    Leptoplana sp.155312191386835163320080602000000004950000003000000

    Turbellaria g. sp.00000000000000060012000108002100000000000

    Nemertini g. sp.30063791340225003764000000001000170045080014001000005003000100200

    Nematoda g. sp.000000000000800000000000000000

    Aricidea claudiae000000000000000000007050402904400000

    Capitella capitata0000004000100000036000000000000000

    Capitomastus minimus000002000300600400200245015001500020000045020037001600

    Eteone picta000000000000000000015000000100000

    Exogone gemmifera000000000000000000050000000000

    Glycera alba000000000000000000000000050000

    Grubea clavata0000000000000003600001000000000000

    Grubea limbata0000000000000000350450000000050100150200

    Harmothoe imbricata000000000000000501005010025000001050000

    Harmothoe reticulata1172171841261192000102000000000000010000000

    Heteromastus filiformis000000000002020125000050008509208301601500360450200160380

    Leiochone clipeata000000000000000000002000000000

    Lisidice nineta000000000000000000050000000000

    Melinna pal0000000000000200000000810270428081205200000

    Microphthalmus shcelcovi00000000000000000000000000000

    Microphthalmus fragilis0000000000000000000000000000800

    Nephthys hombergii0000000000200000000008080905801600000

    Nereis zonata13226105711189510417978500000000002450700094507500000050000

    Nereis succinea551013451628424080120601602020060020014004200505047001850080203010200300200160

    Nerine cirratulus000000000000000005000000000000

    Oridia armandi000000000000000000006005010100000

    Pectinaria coreni0000000000000000000010002000000

    Perinereis cultrifera00000000000000016000000000000000

    Platynereis dumerilii468276768557256412835127504030506000000805054502550167007000000015000100

    Phyllodoce mucosa0000000000000000000030020000000

    Phyllodoce tuberculata14537851119916126340000000000250020030050000000000

    Polycirrus jubatus000000000000000505000000000300000

    Polydora ciliata34403340768126027641201002080701602026050015025505100470024350104001601408024011051504003001850

    Polygordius neapolitanus00000000000000000000000002006018020

    Prionospio cirrifera117001109860140208011080802000502000007501300001102750100200860

    Pygospio elegans0000000000000000200005020080010001000

    Spio filicornis00000000000040120800500050004010000360018005700400

    Sphaerosyllis bulbosa000000000000000050000000000000

    Staurocephalus keffersteini000000000000000000000000045030015050

    Syllides longicirrata0000000000000000000000000250000

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    Syllis prolifera0000000000000003500000000000000

    Oligochaeta g.sp.000001004020604013601020280610013007300250004005900390520309209100300680310

    Athanas nitescens2171171671872340000000000000500000000000

    Ampelisca di0000000000000050000015040101020708050900100

    Amphithoe vail