I N F O M A P B I O R E G I O N S :I N T E R A C T I V E M A P P I N G O F B I O G E O G R A P H I C A L R E G I O N S F R O M S P E C I E S D I S T R I B U T I O N S

Daniel Edler,1, 2, ∗ Thaís Guedes,2, 3, 4 Alexander Zizka,2 Martin Rosvall,1 and Alexandre Antonelli2, 51Integrated Science Lab, Department of Physics, Umeå University, SE-901 87 Umeå, Sweden2University of Gothenburg, Department of Biological and Environmental Sciences, Box 461, SE-405 30 Göteborg,Sweden3Federal University of São Paulo, 09972-270, Diadema, Brazil4Museum of Zoology of University of São Paulo, 04263-000, São Paulo, Brazil5Gothenburg Botanical Garden, Carl Skottsbergs gata 22A, SE-413 19 Gothenburg, Sweden

Biogeographical regions (bioregions) reveal how different sets of species are spatiallygrouped and therefore are important units for conservation, historical biogeography,ecology and evolution. Several methods have been developed to identify bioregions basedon species distribution data rather than expert opinion. One approach successfully appliesnetwork theory to simplify and highlight the underlying structure in species distributions.However, this method lacks tools for simple and efficient analysis. Here we presentInfomap Bioregions, an interactive web application that inputs species distribution dataand generates bioregion maps. Species distributions may be provided as georeferencedpoint occurrences or range maps, and can be of local, regional or global scale. Theapplication uses a novel adaptive resolution method to make best use of often incompletespecies distribution data. The results can be downloaded as vector graphics, shapefilesor in table format. We validate the tool by processing large datasets of publicly availablespecies distribution data of the world’s amphibians using species ranges, and mammalsusing point occurrences. We then calculate the fit between the inferred bioregionsand WWF ecoregions. As examples of applications, researchers can reconstruct ances-tral ranges in historical biogeography or identify indicator species for targeted conservation.

I N T R O D U C T I O N

Biodiversity is not randomly distributed. It is well known thatspecies are grouped in space and patterns of distribution canbe recognized at small and large scales. Depending on the size,source of data and scientific discipline, these broadly used bio-geographical regions have received various related names, buthere we simply refer to them as bioregions (see Vilhena andAntonelli (2015) for a discussion of terminology). In manydisciplines, working with bioregions rather than single speciesis more effective. Conservation biology is a prime example,since protecting bioregions with high levels of biodiversity oruniqueness may help protecting many species from extinction.In historical biogeography, bioregions may be used as oper-ational areas for ancestral range reconstructions in order toestimate how lineages in a phylogeny have evolved their geo-graphical occupancy over time (Goldberg et al., 2011; Matzke,2014; Ree and Smith, 2008). Moreover, since different taxaexhibit different patterns of diversity, distribution, and evolu-tionary history, there is no set of universal bioregions for allcircumstances. Accordingly, the most effective set of biore-gions depends on the particular system under study and re-search question at hand. Therefore, researchers need simple,effective, and flexible tools for mapping relevant species dis-

∗Electronic address: daniel.edler@umu.se

tribution data into bioregions.

While bioinformatic tools can now provide rapid and ac-curate coding of species into predefined areas (TÃupel et al.,2016), choosing the areas in the first place has been a sub-jective procedure without quantitative support. Researchershave therefore developed a suite of algorithms for mappinggrid cell areas into biologically relevant regions (Kozak andWiens, 2006; Kreft and Jetz, 2010; Oliveira et al., 2015), butoften they involve multiple and overly technical steps. As aconsequence, most biogeographical studies still use arbitrarilydefined areas.

To make identification of bioregions simple and effective forany set of species distribution data, we present the web-based,interactive mapping tool Infomap Bioregions. The underlyingmethod clusters bipartite networks that contain both speciesand grid cells. This method was recently shown to outper-form approaches that abstract away the species into speciessimilarities between grid cells in unipartite networks (Vilhenaand Antonelli, 2015). Moreover, the bipartite networks areclustered with the information-theoretic clustering algorithmknown as Infomap (Rosvall and Bergstrom, 2008), which hasbeen acclaimed as the best network clustering algorithm inseveral comparative studies (Aldecoa and Marín, 2013; Lan-cichinetti and Fortunato, 2009). Thanks to its simple and ef-fective design, Infomap Bioregions has wide applications inbiogeography, ecology, evolutionary biology, conservation andrelated disciplines.

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D E S C R I P T I O N

Infomap Bioregions is an interactive web application that iden-tifies taxon-specific bioregions from species distribution data.We first present the application’s workflow (Fig. 1), and thendescribe each step in detail.

Given user-provided species distribution data, the applica-tion first bins the data into geographical grid cells with adaptivespatial resolution. When the data are sparse, the grid size islarge; and when the data are dense, the grid size is small. Thisnovel adaptive resolution offers a considerable advantage overconventional uniform binning when dealing with biodiversitydata, which is unevenly distributed (Maldonado et al., 2015;Meyer et al., 2016).

The binning generates a bipartite network between speciesand grid cells, which is then clustered with the Infomap algo-rithm into bioregions (Edler and Rosvall, 2015). The applica-tion also identifies the most common and the most indicativespecies in each grid cell and bioregion. The results are shownas an interactive map together with supporting tables contain-ing information about each bioregion.

To facilitate the integration of bioregion delimitation andancestral range reconstructions, Infomap Bioregions alsosupports loading a phylogenetic tree, which may be time-calibrated or not. Fitch’s method of maximum parsimony asoriginally described (Fitch, 1971) is implemented to provide aquick estimate of ancestral ranges. Species in the phylogenythat are not present in the distribution dataset are ignored inthe ancestral range reconstruction, and ancestral ranges forthe remaining species are shown with pie charts based on thebioregions identified (see Fig. 3). The bioregions can alsobe exported to allow analyses under alternative methods ofancestral range reconstruction.

Input data

For species distribution data, Infomap Bioregions supportsboth point occurrences and species range maps. Point occur-rences are specified in a text file with either comma-separated(CSV) or tab-separated (TSV) values. The application requiresa header with the column names, and the user must identifywhich columns that should be parsed as name, latitude andlongitude, respectively (Fig. 1a). Range maps are specifiedin the shapefile format, which includes multiple files: a .shpfile for species range polygons, a .dbf file for the attributesof each range polygon and, optionally, a .prj file for projec-tion information. As for point occurrence data, the user mustidentify which attribute to parse as the name of the species.

For phylogenetic data, Infomap Bioregions supports theNEXUS and Newick tree formats.

Output data

The map with bioregions can be exported in .svg and .pngformat. The shapes of bioregions can be exported in .geojsonand shapefile format, and a species presence/absence matrix

Name, Lat, LongSpecies A, 6.9, 1.0Species A, 6.7, 4.7Species A, 1.3, 2.8Species B, 2.3, 2.9Species B, 1.8, 6.4Species B, 4.6, 6.6Species C, 4.7, 5.3Species C, 5.5, 6.8Species C, 6.8, 7.6Species D, 7.4, 6.5Species D, 4.6, 7.4

a) Input data b) Binned data

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Figure 1 Step-by-step illustration of how Infomap Bioregions gener-ates bioregions from species distribution data. Infomap Bioregions:a) inputs comma-separated values for point occurrences, b) adaptivelybins species records into discrete geographical grid cells such that thedata density determines the spatial resolution, c) extracts a bipartitenetwork between species and grid cells, d) clusters the bipartite net-work with the Infomap clustering algorithm, e) visualises the grid cellclusters as bioregions on a zoomable map, f) exports the geograph-ical map in svg or png format, the tables of top occurring and topindicative species for each bioregion in csv format, the species pres-ence/absence matrix for the bioregions in NEXUS format and the ge-ographical information of the bioregions in shapefile or GeoJSONformat.

for the bioregions can be exported in .nexus format for furtheranalyses in e.g. BioGeoBEARS (Matzke, 2013), BayArea(Landis et al., 2013), and many other ancestral reconstructiontools that can handle such files.

Summary tables of the most common and the most indica-tive species for each bioregion can be exported in .csv format,and the tree can be exported in .svg format.

Adaptive resolution

Where data are sparse, single cells can be clustered in distinctbioregions. To avoid providing more detail than the data cansupport, Infomap Bioregions automatically adapts the grid sizeto the amount and spatial distribution of the input data. Thisis done by mapping the input data to a so-called quadtree datastructure, as illustrated in Figure 1b.

The adaptive resolution algorithm uses the quadtree to hi-erarchically partition geographical space into quadratic gridcells of increasingly smaller size by recursively subdividingeach grid cell into four quadrants. When the algorithm reachesthe user-provided maximum cell size (default is 4◦), it aggre-gates the species into grid cells.

To make the resolution adaptive to the density of the data,each grid cell has a user-provided maximum cell capacity (de-fault is 100 species occurrence records). The algorithm re-cursively subdivides all grid cells with more records than themaximum cell capacity until it reaches the user-provided min-imum cell size (default is 1◦). However, if a grid cell after asubdivision contains less species than the user-provided min-

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imum cell capacity (default is 10), the algorithm reverts themost recent subdivision to avoid creating regions with too fewdata points.

With these criteria, Infomap Bioregions can simultaneouslyidentify high-resolution bioregions where data are abundantand low-resolution bioregions where data are sparse, andthereby avoid over- and underfitting across all bioregions.

For point occurrence data, these criteria make the adaptiveresolution straightforward. For range maps, the applicationfirst adds a species record to each grid cell of minimum sizethat intersects with the corresponding species range polygon,and then proceeds with the adaptive binning to satisfy the user-specified criteria.

It is also possible to interactively modify the resolution ofthe bioregions by adjusting the Markov time for the Infomapclustering algorithm (Kheirkhahzadeh et al., 2016). In thisway, the user can tune Infomap to search for bigger or smallerbioregions that are still supported by the data.

Bipartite network

When Infomap Bioregions aggregates the species into geo-graphical grid cells, it forms a bipartite network with speciesand grid cells as the two types of nodes. Each species is con-nected by an unweighted link to each grid cell in which it ispresent. We purposefully avoid weighting the links by thenumber of records, because that would make the results sen-sitive to spatially biased sampling. Instead, we let the densityof species records increase the spatial resolution as describedabove. In this way, dense data generate large networks.

Bioregions and indicator species

Infomap Bioregions clusters the bipartite network with In-fomap for bipartite networks (Kheirkhahzadeh et al., 2016).The resulting clusters contain both grid cells and species, anddefine the bioregions. The software displays the bioregionswith different colours on a map, and provides a table for eachbioregion including summary statistics and species lists. Theapplication lists for grid cells and bioregions both the mostcommon species and the most indicative species with the high-est relative abundance. That is, for species s in grid cell orbioregion r, the indicative score Is|r is defined as the ratio be-tween the frequency fs|r of the species in the region and thefrequency fs of the species in all regions, Is|r = fs|r/ fs. Thusan indicative score of 2 means that a species is twice as fre-quent in the region than in the entire dataset. In the bioregionstables, the most common and indicative species are displayedtogether with charts that show the distribution of those speciesin other bioregions. This information makes it possible to findendemic species, unique or close to unique to a specific biore-gion.

R E S U LT S A N D D I S C U S S I O N

To validate Infomap Bioregions, we applied it to rangemaps of amphibians and point occurrences of terres-trial mammals. For amphibians, we downloaded globaldistribution data as range polygons for 6,069 speciesfrom the International Union for Conservation of Na-ture (IUCN, http://www.iucn.org, downloaded Au-gust 17, 2015, from http://www.iucnredlist.org/technical-documents/spatial-data). For terrestrialmammals, we compiled the global distribution of 5,005 speciesfrom a collection of georeferenced observation records ob-tained through the Global Biodiversity Information Facility(GBIF.org (11th November 2015; GBIF Occurrence Down-load http://doi.org/10.15468/dl.wnjjkc). We cleanedthe mammal dataset using the R functions in the packagespeciesgeocodeR (TÃupel et al., 2016), checking for ob-vious errors such as empty coordinates,terrestrial species re-ported in the sea, and coordinates assigned to country orprovince centroids.

For the resolution, we allowed grid cells to range between4◦ and 2◦ to reflect spatial differences in data density. We usedmaximum cell capacity 100 and set the minimum cell capacityto 5. Below we show the bioregion maps of the amphibiansand mammals, and highlight a few bioregions.

For terrestrial mammals, we downloaded a set of 1000species-level phylogenies of all mammals from (Faurby andSvenning, 2015) and calculated the maximum clade credibil-ity tree using TreeAnnotator in the package BEAST v.1.8.2(Drummond and Rambaut, 2007). We were able to match4,426 species between the phylogeny and the georeferenceddataset.

Amphibians

We identified 87 bioregions of amphibians as illustrated inFigure 2 (see Supplementary Material available on Dryadhttp://dx.doi.org/10.5061/dryad.2s201 for detailedresults). In Table 1, we detail the three most species rich biore-gions and three small bioregions with relatively many species.Most of the species belong to relatively large bioregions, butwe also identified smaller bioregions such as in the Caribbeanwhere island endemics are common and in the tropical Andeswhere species turnover is high and many species are located injust a few cells.

The identified bioregions largely coincide with those foundby Vilhena and Antonelli (2015), except for some differencesdue to the adaptive resolution and its settings. For the Neotrop-ics, our clustering seems to reflect the regionalization proposedby Morrone (2006, 2014) for some sub-regions and provincessuch as the Amazonian subregion, the Parana subregion andthe Chacoan dominion. Infomap Bioregions also successfullyidentified small bioregions, for example in the island of Hispan-iola and in the tropical Andes, which could be particularly con-sidered for conservation. Other examples of relatively small-scale bioregions include the Cape region in South Africa andthe Dahomey gap in West Africa (Fig. 2).

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Figure 2 Bioregion map of the world’s amphibians generated with Infomap Bioregions, using the IUCN species range maps. White areas haveinsufficient data and were excluded from the analysis. The inset shows a zoom-in of Central America, the West Indies and northwestern SouthAmerica, depicting many small bioregions that reflect high turnover of species assemblages and narrow range distributions characteristic forthe region. Table 1 shows information about labelled bioregions.

Table 1 Selected amphibian bioregions. For exact locations, the indices (a)–(f) are displayed on the bioregion map in Figure 2. Bioregions(a)–(c) are the most species rich and (d)–(f) are hand-picked to illustrate how even small bioregions can contain relatively many species.Common names from Encyclopedia of Life at http://eol.org

Location Records Species Cells Most common species (records) Most indicative species (score)(a) South America 42,161 719 167 Trachycephalus venulosus (600) Lithobates palmipes (3.3)

Veined tree frog Amazon River frog(b) Africa 27,267 553 333 Kassina senegalensis (970) Hildebrandtia ornata (2.1)

Senegal running frog African ornate frog(c) Eurasia 13,083 103 313 Rana arvalis (1,547) Triturus cristatus (1.3)

Moor frog Northern crested newt(d) Andes 121 75 1 Pristimantis nervicus (13) Pristimantis nervicus (157)

– –(e) Hispaniola 181 65 4 Hypsiboas heilprini (28) Osteopilus vastus (73)

Los Bracitos tree frog Hispaniola tree frog(f) Cuba 214 61 4 Osteopilus septentrionalis (28) Eleutherodactylus varleyi (73)

Cuban tree frog –

Mammals

We identified 62 bioregions of mammals, which we show to-gether with their phylogenetic tree and ancestral range recon-structions in Figure 3 (see Supplementary Material availableon Dryad http://dx.doi.org/10.5061/dryad.2s201 fordetailed results). Some of the bioregions are very large, reflect-ing major continental-wide differences, while others compriseno more than a few square degrees. For example, we identifiedmore than 10 bioregions for Australia, a landmass known tocontain a high number of species and ecosystems (see Table2).

We acknowledge that the automated cleaning steps de-scribed above for species occurrences are probably not suffi-cient to fully validate the distribution dataset. Careful revisionof specimens and localities by taxonomists, and increased spa-

tial sampling, are some of the time-consuming tasks requiredto produce more reliable datasets (Maldonado et al., 2015;Meyer et al., 2016). As a consequence, our results may beaffected by sampling biases, inaccurate georeferencing, and/orincorrect identifications. These issues prevent us from discern-ing, for example, whether the scattered occurrence of smallbioregions in e.g. Russia is a real biological result or, morelikely, an artefact of the scarce publicly available data for thatregion.

Validation

We further evaluated the performance of Infomap Bioregionsby comparing the bioregions identified for mammals and am-phibians with the widely used WWF ecoregions (Olson et al.,

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Ornithorhynchus anatinus

Megalibgwilia ramsayi

Tachyglossus aculeatus

Zaglossus hacketti

Zaglossus bruijnii

Zaglossus attenboroughi

Zaglossus bartoni

Rhyncholestes raphanurus

Lestoros inca

Caenolestes caniventer

Caenolestes fuliginosus

Caenolestes condorensis

Caenolestes convelatus

Glironia venusta

Caluromysiops irrupta

Caluromys philander

Caluromys derbianus

Caluromys lanatus

Hyladelphys kalinowskii

Tlacuatzin canescens

Chacodelphys formosa

Marmosa rubra

Marmosa mexicana

Marmosa robinsoni

Marmosa xerophila

Marmosa andersoni

Marmosa lepida

Marmosa tyleriana

Marmosa murina

Marmosa quichua

Micoureus paraguayanus

Micoureus demerarae

Micoureus constantiae

Micoureus alstoni

Micoureus regina

Micoureus phaeus

Monodelphis kunsi

Monodelphis emiliae

Monodelphis maraxina

Monodelphis scalops

Monodelphis theresa

Monodelphis osgoodi

Monodelphis umbristriata

Monodelphis reigi

Monodelphis adusta

Monodelphis rubida

Monodelphis unistriata

Monodelphis ronaldi

Monodelphis dimidiata

Monodelphis sorex

Monodelphis americana

Monodelphis iheringi

Monodelphis brevicaudata

Monodelphis domestica

Monodelphis glirina

Monodelphis palliolata

Metachirus nudicaudatus

Chironectes minimus

Lutreolina crassicaudata

Philander frenatus

Philander opossum

Philander mcilhennyi

Philander mondolfii

Philander andersoni

Philander deltae

Didelphis virginiana

Didelphis aurita

Didelphis marsupialis

Didelphis albiventris

Didelphis imperfecta

Didelphis pernigra

Marmosops incanus

Marmosops paulensis

Marmosops noctivagus

Marmosops ocellatus

Marmosops impavidus

Marmosops neblina

Marmosops fuscatus

Marmosops bishopi

Marmosops pinheiroi

Marmosops juninensis

Marmosops creightoni

Marmosops parvidens

Marmosops handleyi

Marmosops invictus

Marmosops cracens

Lestodelphys halli

Thylamys karimii

Thylamys velutinus

Thylamys venustus

Thylamys cinderella

Thylamys sponsorius

Thylamys pusillus

Thylamys macrurus

Thylamys tatei

Thylamys pallidior

Thylamys elegans

Cryptonanus unduaviensis

Cryptonanus chacoensis

Cryptonanus guahybae

Cryptonanus agricolai

Cryptonanus ignitus

Gracilinanus dryas

Gracilinanus emiliae

Gracilinanus marica

Gracilinanus aceramarcae

Gracilinanus agilis

Gracilinanus microtarsus

Dromiciops gliroides

Notoryctes caurinus

Notoryctes typhlops

Chaeropus ecaudatus

Macrotis lagotis

Macrotis leucura

Isoodon macrourus

Isoodon obesulus

Isoodon auratus

Perameles bougainville

Perameles gunnii

Perameles nasuta

Perameles eremiana

Peroryctinae GEN NOW

Peroryctes broadbenti

Peroryctes raffrayana

Peroryctes SP NOW

Microperoryctes longicauda

Microperoryctes papuensis

Microperoryctes murina

Microperoryctes aplini

Echymipera clara

Echymipera echinista

Rhynchomeles prattorum

Echymipera rufescens

Echymipera kalubu

Echymipera davidi

Myrmecobius fasciatus

Thylacinus cynocephalus

Planigale novaeguineae

Planigale maculata

Planigale ingrami

Planigale gilesi

Planigale tenuirostris

Antechinomys laniger

Sminthopsis granulipes

Sminthopsis longicaudata

Ningaui timealeyi

Ningaui yvonneae

Ningaui ridei

Sminthopsis crassicaudata

Sminthopsis bindi

Sminthopsis macroura

Sminthopsis douglasi

Sminthopsis virginiae

Sminthopsis aitkeni

Sminthopsis griseoventer

Sminthopsis ooldea

Sminthopsis psammophila

Sminthopsis youngsoni

Sminthopsis hirtipes

Sminthopsis dolichura

Sminthopsis leucopus

Sminthopsis murina

Sminthopsis gilberti

Sminthopsis archeri

Sminthopsis butleri

Sminthopsis fuliginosus

Phascogale calura

Phascogale tapoatafa

Phascogale pirata

Murexia habbema

Murexia rothschildi

Murexia melanurus

Murexia longicaudata

Murexia naso

Antechinus swainsonii

Antechinus minimus

Antechinus subtropicus

Antechinus godmani

Antechinus stuartii

Antechinus adustus

Antechinus flavipes

Antechinus bellus

Antechinus leo

Antechinus agilis

Dasyuroides byrnei

Dasykaluta rosamondae

Dasycercus cristicauda

Dasycercus blythi

Pseudantechinus bilarni

Pseudantechinus woolleyae

Pseudantechinus ningbing

Pseudantechinus macdonnellensis

Pseudantechinus roryi

Pseudantechinus mimulus

Parantechinus apicalis

Myoictis melas

Myoictis wallacei

Myoictis wavicus

Myoictis leucura

Neophascogale lorentzii

Phascolosorex dorsalis

Phascolosorex doriae

Sarcophilus harrisii

Dasyurus hallucatus

Dasyurus maculatus

Dasyurus albopunctatus

Dasyurus viverrinus

Dasyurus geoffroii

Dasyurus spartacus

Phascolarctos cinereus

Thylacoleo carnifex

Palorchestes azeal

Diprotodon optatum

Maokopia ronaldi

Zygomaturus trilobus

Vombatus ursinus

Vombatus hacketti

Phascolomys medius

Phascolonus gigas

Lasiorhinus krefftii

Lasiorhinus latifrons

Burramys parvus

Cercartetus caudatus

Cercartetus lepidus

Cercartetus nanus

Cercartetus concinnus

Wyulda squamicaudata

Trichosurus vulpecula

Trichosurus caninus

Trichosurus cunninghami

Strigocuscus celebensis

Ailurops ursinus

Ailurops melanotis

Spilocuscus papuensis

Spilocuscus rufoniger

Spilocuscus wilsoni

Spilocuscus maculatus

Spilocuscus kraemeri

Strigocuscus pelengensis

Phalanger matanim

Phalanger ornatus

Phalanger lullulae

Phalanger gymnotis

Phalanger sericeus

Phalanger vestitus

Phalanger carmelitae

Phalanger intercastellanus

Phalanger alexandrae

Phalanger orientalis

Phalanger matabiru

Phalanger mimicus

Phalanger rothschildi

Acrobates pygmaeus

Distoechurus pennatus

Tarsipes rostratus

Gymnobelideus leadbeateri

Dactylonax palpator

Dactylopsila trivirgata

Dactylopsila tatei

Dactylopsila megalura

Dactylopsila kambuayai

Petaurus australis

Petaurus abidi

Petaurus breviceps

Petaurus biacensis

Petaurus gracilis

Petaurus norfolcensis

Pseudochirops archeri

Petropseudes dahli

Pseudochirops corinnae

Pseudochirops cupreus

Pseudochirops albertisii

Pseudochirops coronatus

Hemibelideus lemuroides

Petauroides volans

Petauroides ayamaruensis

Pseudocheirus peregrinus

Pseudocheirus occidentalis

Pseudochirulus herbertensis

Pseudochirulus cinereus

Pseudochirulus canescens

Pseudochirulus schlegeli

Pseudochirulus forbesi

Pseudochirulus larvatus

Pseudochirulus caroli

Pseudochirulus mayeri

Hypsiprymnodon moschatus

Potorous tridactylus

Potorous platyops

Potorous gilbertii

Potorous longipes

Caloprymnus campestris

Borungaboodie hatcheri

Aepyprymnus rufescens

Propleopus oscillans

Bettongia penicillata

Bettongia gaimardi

Bettongia tropica

Bettongia lesueur

Bettongia pusilla

Sthenurus andersoni

Sthenurus atlas

Sthenurus stirlingi

Sthenurus tindalei

Metasthenurus newtonae

Simosthenurus occidentalis

Procoptodon browneorum

Procoptodon goliah

Procoptodon gilli

Lagostrophus fasciatus

Troposodon minor

Onychogalea fraenata

Onychogalea unguifera

Onychogalea lunata

Setonix brachyurus

Lagorchestes asomatus

Lagorchestes hirsutus

Lagorchestes conspicillatus

Lagorchestes leporides

Macropus fuliginosus

Macropus giganteus

Macropus ferragus

Wallabia bicolor

Wallabia kitcheneri

Macropus bernardus

Macropus rufus

Macropus antilopinus

Macropus robustus

Macropus irma

Macropus rufogriseus

Macropus parryi

Macropus parma

Macropus dorsalis

Macropus agilis

Macropus eugenii

Macropus greyi

Dorcopsulus vanheurni

Dorcopsulus macleayi

Dorcopsis luctuosa

Dorcopsis muelleri

Dorcopsis atrata

Dorcopsis hageni

Protemnodon nombe

Protemnodon roechus

Protemnodon hopei

Protemnodon tumbuna

Protemnodon anak

Protemnodon brehus

Thylogale billardierii

Thylogale thetis

Thylogale stigmatica

Thylogale christenseni

Thylogale lanatus

Thylogale browni

Thylogale calabyi

Thylogale brunii

Dendrolagus inustus

Dendrolagus spadix

Dendrolagus matschiei

Dendrolagus pulcherrimus

Dendrolagus goodfellowi

Dendrolagus mbaiso

Dendrolagus lumholtzi

Dendrolagus bennettianus

Dendrolagus ursinus

Dendrolagus dorianus

Dendrolagus mayri

Dendrolagus notatus

Dendrolagus stellarum

Dendrolagus scottae

Petrogale persephone

Petrogale burbidgei

Petrogale concinna

Petrogale brachyotis

Petrogale xanthopus

Petrogale coenensis

Petrogale lateralis

Petrogale godmani

Petrogale rothschildi

Petrogale purpureicollis

Petrogale herberti

Petrogale penicillata

Petrogale assimilis

Petrogale sharmani

Petrogale inornata

Petrogale mareeba

Mixotoxodon larensis

Toxodon platensis

Trigonodops lopesi

Dugong dugon

Hydrodamalis gigas

Trichechus senegalensis

Trichechus inunguis

Trichechus manatus

Procavia capensis

Heterohyrax brucei

Dendrohyrax dorsalis

Dendrohyrax arboreus

Dendrohyrax validus

Mammut americanum

Cuvieronius hyodon

Stegomastodon platensis

Stegomastodon waringi

Stegodon florensis

Stegodon orientalis

Stegodon trigonocephalus

Mammuthus primigenius

Mammuthus columbi

Mammuthus exilis

Loxodonta africana

Elephas iolensis

Elephas maximus

Elephas mnaidriensis

Elephas naumanni

Elephas namadicus

Elephas cypriotes

Elephas tiliensis

Elephas antiquus

Orycteropus afer

Plesiorycteropus madagascariensis

Rhynchocyon chrysopygus

Rhynchocyon udzungwensis

Rhynchocyon petersi

Rhynchocyon cirnei

Macroscelides proboscideus

Elephantulus rozeti

Petrodromus tetradactylus

Elephantulus revoili

Elephantulus fuscipes

Elephantulus fuscus

Elephantulus myurus

Elephantulus pilicaudus

Elephantulus edwardii

Elephantulus rupestris

Elephantulus intufi

Elephantulus rufescens

Elephantulus brachyrhynchus

Chrysospalax trevelyani

Chrysospalax villosus

Calcochloris obtusirostris

Calcochloris leucorhinus

Calcochloris tytonis

Eremitalpa granti

Cryptochloris wintoni

Cryptochloris zyli

Chrysochloris stuhlmanni

Chrysochloris asiatica

Chrysochloris visagiei

Chlorotalpa duthieae

Chlorotalpa sclateri

Carpitalpa arendsi

Neamblysomus gunningi

Neamblysomus julianae

Amblysomus marleyi

Amblysomus corriae

Amblysomus hottentotus

Amblysomus septentrionalis

Amblysomus robustus

Potamogale velox

Micropotamogale lamottei

Micropotamogale ruwenzorii

Setifer setosus

Echinops telfairi

Tenrec ecaudatus

Hemicentetes semispinosus

Hemicentetes nigriceps

Geogale aurita

Oryzorictes hova

Oryzorictes tetradactylus

Limnogale mergulus

Microgale soricoides

Microgale macpheei

Microgale nasoloi

Microgale parvula

Microgale drouhardi

Microgale cowani

Microgale brevicaudata

Microgale dobsoni

Microgale pusilla

Microgale gymnorhyncha

Microgale gracilis

Microgale taiva

Microgale majori

Microgale monticola

Microgale principula

Microgale jobihely

Microgale fotsifotsy

Microgale talazaci

Microgale dryas

Microgale thomasi

Microgale longicaudata

Microgale jenkinsae

Cyclopes didactylus

Myrmecophaga tridactyla

Tamandua mexicana

Tamandua tetradactyla

Bradypus torquatus

Bradypus tridactylus

Bradypus variegatus

Bradypus pygmaeus

Nothrotherium maquinense

Nothrotheriops shastense

Eremotherium laurillardi

Megatherium americanum

Megatherium tarijense

Scelidotherium leptocephalum

Catonyx cuvieri

Scelidodon chiliensis

Valgipes deformis

Paramylodon harlani

Mylodon darwinii

Glossotherium robustum

Lestodon armatus

Choloepus hoffmanni

Choloepus didactylus

Neocnus comes

Neocnus dousman

Neocnus toupiti

Megalonyx jeffersonii

Diabolotherium nordenskioldi

Acratocnus odontrigonus

Acratocnus ye

Parocnus browni

Parocnus serus

Megalocnus zile

Megalocnus rodens

Priodontes maximus

Tolypeutes tricinctus

Tolypeutes matacus

Cabassous chacoensis

Cabassous centralis

Cabassous unicinctus

Cabassous tatouay

Chlamyphorus truncatus

Chlamyphorus retusus

Euphractus sexcinctus

Zaedyus pichiy

Eutatus seguini

Chaetophractus villosus

Chaetophractus vellerosus

Chaetophractus nationi

Propraopus sulcatus

Propraopus punctatus

Dasypus kappleri

Dasypus novemcinctus

Dasypus yepesi

Dasypus bellus

Dasypus sabanicola

Dasypus hybridus

Dasypus septemcinctus

Dasypus pilosus

Pampatherium humboldti

Pampatherium typum

Holmesina septentrionalis

Holmesina occidentalis

Holmesina paulacoutoi

Neosclerocalyptus paskoenis

Panochthus tuberculatus

Hoplophorus euphractus

Glyptodon clavipes

Glyptodon reticulatus

Doedicurus clavicaudatus

Neuryurus trabeculatus

Glyptotherium cylindricum

Glyptotherium floridanum

Solenodon paradoxus

Solenodon cubanus

Solenodon marcanoi

Nesophontes edithae

Nesophontes SP A

Nesophontes SP B

Nesophontes paramicrus

Nesophontes hypomicrus

Nesophontes major

Nesophontes zamicrus

Nesophontes micrus

Neurotrichus gibbsii

Scaptonyx fusicaudus

Scapanulus oweni

Parascalops breweri

Scalopus aquaticus

Scapanus latimanus

Scapanus orarius

Scapanus townsendii

Urotrichus talpoides

Dymecodon pilirostris

Galemys pyrenaicus

Desmana moschata

Condylura cristata

Uropsilus soricipes

Uropsilus andersoni

Uropsilus investigator

Uropsilus gracilis

Talpa altaica

Talpa levantis

Talpa stankovici

Talpa caucasica

Talpa romana

Talpa caeca

Talpa europaea

Talpa occidentalis

Talpa davidiana

Euroscaptor mizura

Euroscaptor micrura

Scaptochirus moschatus

Euroscaptor longirostris

Euroscaptor parvidens

Euroscaptor grandis

Euroscaptor klossi

Parascaptor leucura

Mogera uchidai

Mogera insularis

Mogera tokudae

Mogera etigo

Mogera imaizumii

Mogera wogura

Mogera robusta

Echinosorex gymnura

Podogymnura truei

Podogymnura aureospinula

Neohylomys hainanensis

Neotetracus sinensis

Hylomys parvus

Hylomys suillus

Hylomys megalotis

Hemiechinus auritus

Hemiechinus collaris

Mesechinus hughi

Mesechinus dauuricus

Paraechinus hypomelas

Paraechinus aethiopicus

Paraechinus micropus

Paraechinus nudiventris

Atelerix frontalis

Atelerix algirus

Atelerix albiventris

Atelerix sclateri

Erinaceus europaeus

Erinaceus amurensis

Erinaceus concolor

Erinaceus roumanicus

Anourosorex squamipes

Anourosorex assamensis

Anourosorex schmidi

Anourosorex yamashinai

Megasorex gigas

Notiosorex villai

Notiosorex evotis

Notiosorex cockrumi

Notiosorex crawfordi

Neomys fodiens

Neomys teres

Neomys anomalus

Episoriculus fumidus

Chodsigoa sodalis

Chodsigoa caovansunga

Chodsigoa parca

Chodsigoa lamula

Chodsigoa parva

Chodsigoa salenskii

Chodsigoa hypsibia

Chodsigoa smithii

Soriculus nigrescens

Nesiotites hidalgo

Nesiotites similis

Episoriculus macrurus

Episoriculus leucops

Episoriculus caudatus

Nectogale elegans

Chimarrogale hantu

Chimarrogale phaeura

Chimarrogale sumatrana

Chimarrogale styani

Chimarrogale himalayica

Chimarrogale platycephalus

Blarinella wardi

Blarinella griselda

Blarinella quadraticauda

Blarina hylophaga

Blarina carolinensis

Blarina brevicauda

Cryptotis gracilis

Cryptotis endersi

Cryptotis magna

Cryptotis squamipes

Cryptotis montivaga

Cryptotis meridensis

Cryptotis peregrina

Cryptotis mexicana

Cryptotis phillipsii

Cryptotis nelsoni

Cryptotis obscura

Cryptotis brachyonyx

Cryptotis colombiana

Cryptotis goodwini

Cryptotis thomasi

Cryptotis equatoris

Cryptotis medellinia

Cryptotis merriami

Cryptotis mayensis

Cryptotis nigrescens

Cryptotis mera

Cryptotis tropicalis

Cryptotis parva

Cryptotis orophila

Cryptotis hondurensis

Cryptotis griseoventris

Cryptotis goldmani

Cryptotis peruviensis

Cryptotis alticola

Cryptotis tamensis

Sorex alpinus

Sorex cylindricauda

Sorex excelsus

Sorex bedfordiae

Sorex mirabilis

Sorex raddei

Sorex roboratus

Sorex samniticus

Sorex arcticus

Sorex maritimensis

Sorex tundrensis

Sorex asper

Sorex daphaenodon

Sorex coronatus

Sorex granarius

Sorex araneus

Sorex satunini

Sorex antinorii

Sorex arunchi

Sorex gracillimus

Sorex shinto

Sorex caecutiens

Sorex cansulus

Sorex unguiculatus

Sorex milleri

Sorex isodon

Sorex sinalis

Sorex yukonicus

Sorex minutissimus

Sorex hosonoi

Sorex minutus

Sorex volnuchini

Sorex kozlovi

Sorex thibetanus

Sorex ventralis

Sorex buchariensis

Sorex planiceps

Sorex arizonae

Sorex trowbridgii

Sorex merriami

Sorex veraecrucis

Sorex stizodon

Sorex mediopua

Sorex saussurei

Sorex veraepacis

Sorex ixtlanensis

Sorex dispar

Sorex emarginatus

Sorex fumeus

Sorex tenellus

Sorex macrodon

Sorex vagrans

Sorex ornatus

Sorex monticolus

Sorex neomexicanus

Sorex sonomae

Sorex alaskanus

Sorex palustris

Sorex sclateri

Sorex bendirii

Sorex pacificus

Sorex bairdi

Sorex leucogaster

Sorex rohweri

Sorex hoyi

Sorex cinereus

Sorex longirostris

Sorex oreopolus

Sorex orizabae

Sorex camtschatica

Sorex jacksoni

Sorex nanus

Sorex ugyunak

Sorex portenkoi

Sorex pribilofensis

Sorex lyelli

Sorex haydeni

Sorex preblei

Surdisorex norae

Surdisorex polulus

Congosorex verheyeni

Congosorex phillipsorum

Congosorex polli

Myosorex zinki

Myosorex schalleri

Myosorex kihaulei

Myosorex geata

Myosorex blarina

Myosorex rumpii

Myosorex longicaudatus

Myosorex tenuis

Myosorex sclateri

Myosorex okuensis

Myosorex varius

Myosorex eisentrauti

Myosorex cafer

Myosorex babaulti

Feroculus feroculus

Sylvisorex lunaris

Sylvisorex johnstoni

Sylvisorex vulcanorum

Sylvisorex howelli

Suncus remyi

Suncus infinitesimus

Suncus varilla

Suncus zeylanicus

Suncus aequatorius

Suncus lixus

Suncus ater

Ruwenzorisorex suncoides

Suncus megalura

Scutisorex somereni

Sylvisorex konganensis

Sylvisorex morio

Sylvisorex isabellae

Sylvisorex granti

Sylvisorex camerunensis

Sylvisorex oriundus

Sylvisorex ollula

Sylvisorex pluvialis

Solisorex pearsoni

Suncus malayanus

Suncus madagascariensis

Suncus etruscus

Suncus fellowesgordoni

Suncus hosei

Suncus mertensi

Suncus dayi

Suncus stoliczkanus

Suncus murinus

Suncus montanus

Crocidura pachyura

Crocidura russula

Crocidura katinka

Crocidura nanilla

Crocidura macarthuri

Paracrocidura schoutedeni

Paracrocidura maxima

Paracrocidura graueri

Crocidura douceti

Crocidura muricauda

Crocidura nimbae

Crocidura wimmeri

Crocidura parvipes

Crocidura lamottei

Crocidura vorax

Crocidura fulvastra

Crocidura viaria

Crocidura goliath

Crocidura olivieri

Crocidura mutesae

Crocidura kivuana

Crocidura nigrofusca

Crocidura flavescens

Crocidura yankariensis

Crocidura ansellorum

Crocidura usambarae

Crocidura hirta

Crocidura andamanensis

Crocidura greenwoodi

Crocidura tansaniana

Crocidura xantippe

Crocidura hutanis

Crocidura zimmeri

Crocidura caliginea

Crocidura crenata

Crocidura ludia

Crocidura polia

Crocidura grassei

Crocidura telfordi

Crocidura fuscomurina

Crocidura planiceps

Crocidura pitmani

Crocidura maquassiensis

Crocidura montis

Crocidura lusitania

Crocidura allex

Crocidura jouvenetae

Crocidura crossei

Crocidura arabica

Crocidura roosevelti

Crocidura jacksoni

Crocidura dolichura

Crocidura latona

Crocidura cyanea

Crocidura gracilipes

Crocidura zaphiri

Crocidura silacea

Crocidura mariquensis

Crocidura baluensis

Crocidura desperata

Crocidura voi

Crocidura fischeri

Crocidura maurisca

Crocidura denti

Crocidura congobelgica

Crocidura smithii

Crocidura lea

Crocidura ultima

Crocidura macowi

Crocidura virgata

Crocidura bottegoides

Crocidura hildegardeae

Crocidura hilliana

Crocidura religiosa

Crocidura grandiceps

Crocidura theresae

Crocidura foxi

Crocidura longipes

Crocidura batesi

Crocidura turba

Crocidura tarella

Crocidura nigeriae

Crocidura buettikoferi

Crocidura attila

Crocidura poensis

Crocidura phaeura

Crocidura harenna

Crocidura musseri

Crocidura rhoditis

Crocidura elongata

Crocidura levicula

Crocidura leucodon

Crocidura zimmermanni

Crocidura pasha

Crocidura ramona

Crocidura sicula

Crocidura canariensis

Crocidura tarfayensis

Crocidura susiana

Crocidura obscurior

Crocidura bottegi

Crocidura lanosa

Crocidura monax

Crocidura littoralis

Crocidura stenocephala

Crocidura macmillani

Crocidura grandis

Crocidura fumosa

Crocidura selina

Crocidura luna

Crocidura vosmaeri

Crocidura glassi

Crocidura nana

Crocidura thalia

Crocidura somalica

Crocidura dhofarensis

Crocidura baileyi

Crocidura lucina

Crocidura pullata

Crocidura arispa

Crocidura armenica

Crocidura serezkyensis

Crocidura pergrisea

Crocidura zarudnyi

Crocidura shantungensis

Crocidura aleksandrisi

Crocidura caspica

Crocidura sibirica

Crocidura floweri

Crocidura suaveolens

Crocidura gmelini

Diplomesodon pulchellum

Crocidura orii

Crocidura eisentrauti

Crocidura niobe

Crocidura horsfieldii

Crocidura cinderella

Crocidura hikmiya

Crocidura manengubae

Crocidura miya

Crocidura nigricans

Crocidura erica

Crocidura tanakae

Crocidura attenuata

Crocidura trichura

Crocidura paradoxura

Crocidura raineyi

Crocidura hispida

Crocidura indochinensis

Crocidura wuchihensis

Crocidura fuliginosa

Crocidura tenuis

Crocidura dsinezumi

Crocidura lasiura

Crocidura whitakeri

Crocidura rapax

Crocidura nicobarica

Crocidura jenkinsi

Crocidura watasei

Crocidura maxi

Crocidura monticola

Crocidura mindorus

Crocidura foetida

Crocidura nigripes

Crocidura palawanensis

Crocidura grayi

Crocidura negrina

Crocidura beatus

Crocidura negligens

Crocidura malayana

Crocidura elgonius

Crocidura brunnea

Crocidura orientalis

Crocidura lepidura

Crocidura thomensis

Crocidura beccarii

Crocidura picea

Equus ovodovi

Equus hydruntinus

Hippidion principale

Hippidion devillei

Equus ferus

Equus semiplicatus

Equus africanus

Equus zebra

Equus grevyi

Equus quagga

Equus hemionus

Equus kiang

Tapirus indicus

Tapirus augustus

Tapirus bairdii

Tapirus copei

Tapirus veroensis

Tapirus rondoniensis

Tapirus terrestris

Tapirus pinchaque

Elasmotherium sibiricum

Diceros bicornis

Ceratotherium simum

Rhinoceros sondaicus

Rhinoceros unicornis

Dicerorhinus sumatrensis

Coelodonta antiquitatis

Stephanorhinus hemitoechus

Stephanorhinus kirchbergensis

Smutsia temminckii

Smutsia gigantea

Uromanis tetradactyla

Phataginus tricuspis

Manis culionensis

Manis javanica

Manis paleojavanica

Manis pentadactyla

Manis crassicaudata

Nandinia binotata

Prionodon pardicolor

Prionodon linsang

Homotherium latidens

Homotherium serum

Smilodon fatalis

Smilodon populator

Neofelis nebulosa

Neofelis diardi

Panthera tigris

Panthera uncia

Panthera pardus

Panthera onca

Panthera leo

Panthera spelaea

Panthera atrox

Leptailurus serval

Caracal caracal

Caracal aurata

Pardofelis marmorata

Pardofelis temminckii

Pardofelis badia

Leopardus pardalis

Leopardus wiedii

Leopardus amnicola

Leopardus jacobita

Leopardus colocolo

Leopardus tigrinus

Leopardus guigna

Leopardus geoffroyi

Acinonyx jubatus

Puma yagouaroundi

Puma concolor

Miracinonyx trumani

Lynx rufus

Lynx pardinus

Lynx canadensis

Lynx lynx

Felis chaus

Felis nigripes

Felis margarita

Felis silvestris

Otocolobus manul

Prionailurus rubiginosus

Prionailurus planiceps

Prionailurus bengalensis

Prionailurus viverrinus

Cynogale bennettii

Chrotogale owstoni

Hemigalus derbyanus

Diplogale hosei

Arctogalidia trivirgata

Arctictis binturong

Paguma larvata

Macrogalidia musschenbroekii

Paradoxurus hermaphroditus

Paradoxurus jerdoni

Paradoxurus zeylonensis

Viverricula indica

Civettictis civetta

Viverra tangalunga

Viverra zibetha

Viverra megaspila

Viverra civettina

Poiana richardsonii

Poiana leightoni

Genetta cristata

Genetta thierryi

Genetta abyssinica

Genetta servalina

Genetta victoriae

Genetta johnstoni

Genetta piscivora

Genetta genetta

Genetta tigrina

Genetta angolensis

Genetta maculata

Genetta bourloni

Genetta poensis

Genetta pardina

Crocuta crocuta

Proteles cristata

Hyaena hyaena

Hyaena brunnea

Fossa fossana

Eupleres goudotii

Cryptoprocta ferox

Cryptoprocta spelea

Galidia elegans

Salanoia concolor

Mungotictis decemlineata

Galidictis fasciata

Galidictis grandidieri

Liberiictis kuhni

Mungos mungo

Mungos gambianus

Suricata suricatta

Dologale dybowskii

Helogale parvula

Helogale hirtula

Crossarchus alexandri

Crossarchus ansorgei

Crossarchus obscurus

Crossarchus platycephalus

Herpestes ichneumon

Herpestes pulverulentus

Herpestes flavescens

Herpestes sanguineus

Herpestes ochraceus

Ichneumia albicauda

Rhynchogale melleri

Paracynictis selousi

Cynictis penicillata

Bdeogale nigripes

Bdeogale jacksoni

Bdeogale crassicauda

Bdeogale omnivora

Atilax paludinosus

Herpestes naso

Herpestes smithii

Herpestes brachyurus

Herpestes urva

Herpestes vitticollis

Herpestes fuscus

Herpestes edwardsii

Herpestes javanicus

Herpestes semitorquatus

Urocyon cinereoargenteus

Urocyon littoralis

Otocyon megalotis

Nyctereutes procyonoides

Vulpes zerda

Vulpes cana

Vulpes chama

Vulpes pallida

Vulpes velox

Vulpes macrotis

Alopex lagopus

Vulpes vulpes

Vulpes rueppellii

Vulpes bengalensis

Vulpes corsac

Vulpes ferrilata

Canis adustus

Canis mesomelas

Lycaon pictus

Cuon alpinus

Cynotherium sardous

Canis dirus

Canis lupus

Canis rufus

Canis aureus

Canis simensis

Canis latrans

Speothos venaticus

Chrysocyon brachyurus

Dusicyon australis

Dusicyon avus

Atelocynus microtis

Protocyon troglodytes

Cerdocyon thous

Theriodictis tarijensis

Pseudalopex culpaeus

Pseudalopex fulvipes

Pseudalopex sechurae

Pseudalopex vetulus

Pseudalopex griseus

Pseudalopex gymnocercus

Ailuropoda melanoleuca

Tremarctos ornatus

Tremarctos floridanus

Arctodus simus

Arctotherium tarijense

Arctotherium wingei

Helarctos malayanus

Melursus ursinus

Ursus thibetanus

Ursus americanus

Ursus spelaeus

Ursus maritimus

Ursus arctos

Odobenus rosmarus

Callorhinus ursinus

Otaria flavescens

Arctocephalus pusillus

Arctocephalus tropicalis

Eumetopias jubatus

Zalophus japonicus

Zalophus californianus

Zalophus wollebaeki

Arctocephalus gazella

Phocarctos hookeri

Neophoca cinerea

Arctocephalus philippii

Arctocephalus townsendi

Arctocephalus forsteri

Arctocephalus australis

Arctocephalus galapagoensis

Monachus monachus

Monachus schauinslandi

Monachus tropicalis

Mirounga angustirostris

Mirounga leonina

Ommatophoca rossii

Lobodon carcinophaga

Hydrurga leptonyx

Leptonychotes weddellii

Erignathus barbatus

Cystophora cristata

Pagophilus groenlandicus

Histriophoca fasciata

Phoca largha

Phoca vitulina

Halichoerus grypus

Pusa sibirica

Pusa hispida

Pusa caspica

Mydaus marchei

Mydaus javanensis

Brachyprotoma obtusata

Conepatus leuconotus

Conepatus chinga

Conepatus humboldtii

Conepatus semistriatus

Mephitis mephitis

Mephitis macroura

Spilogale pygmaea

Spilogale gracilis

Spilogale putorius

Spilogale angustifrons

Ailurus fulgens

Potos flavus

Bassariscus astutus

Bassariscus sumichrasti

Procyon lotor

Procyon cancrivorus

Procyon pygmaeus

Nasua nasua

Nasua narica

Nasuella olivacea

Bassaricyon beddardi

Bassaricyon pauli

Bassaricyon lasius

Bassaricyon alleni

Bassaricyon gabbii

Taxidea taxus

Mellivora capensis

Arctonyx collaris

Meles meles

Meles leucurus

Meles anakuma

Eira barbara

Martes pennanti

Gulo gulo

Martes flavigula

Martes gwatkinsii

Martes foina

Martes melampus

Martes zibellina

Martes martes

Martes americana

Melogale orientalis

Melogale everetti

Melogale moschata

Melogale personata

Lyncodon patagonicus

Galictis vittata

Galictis cuja

Vormela peregusna

Ictonyx libyca

Ictonyx striatus

Poecilogale albinucha

Pteronura brasiliensis

Lontra canadensis

Lontra longicaudis

Lontra provocax

Lontra felina

Enhydra lutris

Lutra maculicollis

Lutra lutra

Lutra sumatrana

Aonyx cinerea

Lutrogale perspicillata

Aonyx capensis

Aonyx congicus

Neovison vison

Neovison macrodon

Mustela frenata

Mustela felipei

Mustela africana

Mustela nudipes

Mustela strigidorsa

Mustela kathiah

Mustela erminea

Mustela altaica

Mustela nivalis

Mustela subpalmata

Mustela lutreolina

Mustela itatsi

Mustela sibirica

Mustela putorius

Mustela eversmanii

Mustela nigripes

Mustela lutreola

Neolicaphrium recens

Macrauchenia patachonica

Xenorhinotherium bahiense

Camelus dromedarius

Camelus ferus

Palaeolama weddeli

Palaeolama major

Palaeolama mirifica

Camelops hesternus

Hemiauchenia macrocephala

Hemiauchenia paradoxa

Lama guanicoe

Vicugna vicugna

Tayassu pecari

Platygonus compressus

Mylohyus nasutus

Catagonus wagneri

Catagonus stenocephalus

Pecari tajacu

Pecari maximus

Babyrousa togeanensis

Babyrousa babyrussa

Babyrousa celebensis

Potamochoerus larvatus

Potamochoerus porcus

Hylochoerus meinertzhageni

Kolpochoerus majus

Metridiochoerus compactus

Phacochoerus aethiopicus

Phacochoerus africanus

Porcula salvania

Sus scrofa

Sus celebensis

Sus bucculentus

Sus philippensis

Sus cebifrons

Sus barbatus

Sus ahoenobarbus

Sus oliveri

Sus verrucosus

Sus brachgnathus

Choeropsis liberiensis

Hexaprotodon sivalensis

Hippopotamus lemerlei

Hippopotamus amphibius

Phanourios minutes

Hippopotamus madagascariensis

Eubalaena australis

Balaena mysticetus

Eubalaena japonica

Eubalaena glacialis

Caperea marginata

Eschrichtius robustus

Balaenoptera bonaerensis

Balaenoptera acutorostrata

Balaenoptera musculus

Megaptera novaeangliae

Balaenoptera physalus

Balaenoptera edeni

Balaenoptera borealis

Balaenoptera omurai

Physeter macrocephalus

Kogia breviceps

Kogia sima

Platanista gangetica

Berardius bairdii

Berardius arnuxii

Tasmacetus shepherdi

Ziphius cavirostris

Hyperoodon planifrons

Hyperoodon ampullatus

Mesoplodon bidens

Mesoplodon layardii

Mesoplodon bowdoini

Mesoplodon carlhubbsi

Mesoplodon mirus

Mesoplodon europaeus

Mesoplodon traversii

Mesoplodon ginkgodens

Mesoplodon hectori

Indopacetus pacificus

Mesoplodon grayi

Mesoplodon perrini

Mesoplodon peruvianus

Mesoplodon densirostris

Mesoplodon stejnegeri

Lipotes vexillifer

Inia geoffrensis

Pontoporia blainvillei

Delphinapterus leucas

Monodon monoceros

Neophocaena phocaenoides

Phocoenoides dalli

Phocoena phocoena

Phocoena sinus

Phocoena spinipinnis

Phocoena dioptrica

Lagenorhynchus acutus

Lissodelphis borealis

Lissodelphis peronii

Lagenorhynchus obliquidens

Lagenorhynchus obscurus

Lagenorhynchus cruciger

Lagenorhynchus australis

Cephalorhynchus heavisidii

Cephalorhynchus hectori

Cephalorhynchus eutropia

Cephalorhynchus commersonii

Lagenorhynchus albirostris

Sotalia fluviatilis

Orcinus orca

Orcaella brevirostris

Orcaella heinsohni

Steno bredanensis

Pseudorca crassidens

Grampus griseus

Feresa attenuata

Peponocephala electra

Globicephala melas

Globicephala macrorhynchus

Lagenodelphis hosei

Stenella attenuata

Sousa chinensis

Sousa teuszii

Stenella longirostris

Delphinus capensis

Delphinus delphis

Stenella coeruleoalba

Stenella clymene

Tursiops aduncus

Stenella frontalis

Tursiops truncatus

Moschiola indica

Moschiola kathygre

Moschiola meminna

Hyemoschus aquaticus

Tragulus williamsoni

Tragulus javanicus

Tragulus kanchil

Tragulus versicolor

Tragulus napu

Tragulus nigricans

Giraffa camelopardalis

Okapia johnstoni

Antilocapra americana

Capromeryx minor

Stockoceros conklingi

Tetrameryx shuleri

Moschus anhuiensis

Moschus berezovskii

Moschus fuscus

Moschus chrysogaster

Moschus moschiferus

Moschus leucogaster

Moschus cupreus

Hydropotes inermis

Capreolus capreolus

Capreolus miyakoensis

Capreolus pygargus

Alces scotti

Alces alces

Alces americanus

Rangifer tarandus

Agalmaceros blicki

Morenelaphus brachyceros

Morenelaphus lujanensis

Pudu puda

Pudu mephistophiles

Mazama nemorivaga

Mazama gouazoubira

Blastocerus dichotomus

Ozotoceros bezoarticus

Hippocamelus antisensis

Hippocamelus bisulcus

Paraceros fragilis

Navahoceros fricki

Odocoileus virginianus

Odocoileus hemionus

Sangamona fugitiva

Antifer ultra

Mazama rufina

Mazama chunyi

Mazama bororo

Mazama nana

Mazama bricenii

Mazama temama

Mazama americana

Mazama pandora

Elaphodus cephalophus

Dicroceros sp

Muntiacus reevesi

Muntiacus atherodes

Muntiacus vaginalis

Muntiacus muntjak

Muntiacus montanus

Muntiacus vuquangensis

Muntiacus truongsonensis

Muntiacus putaoensis

Muntiacus crinifrons

Muntiacus gongshanensis

Muntiacus feae

Muntiacus rooseveltorum

Megaceroides algericus

Sinomegaceros yabei

Sinomegaceros ordosianus

Megaloceros giganteus

Praemegaceros cazioti

Candiacervus dorothensis

Candiacervus SpII

Candiacervus cretensis

Candiacervus rethymnensis

Candiacervus ropalophorus

Candiacervus major

Dama dama

Dama mesopotamica

Rucervus eldii

Rucervus schomburgki

Rucervus duvaucelii

Axis axis

Axis calamianensis

Axis porcinus

Axis kuhlii

Elaphurus davidianus

Rusa unicolor

Rusa timorensis

Rusa alfredi

Rusa marianna

Przewalskium albirostris

Cervus elaphus

Cervus nippon

Cervus astylodon

Tetracerus quadricornis

Boselaphus tragocamelus

Tragelaphus angasii

Tragelaphus derbianus

Tragelaphus oryx

Tragelaphus strepsiceros

Tragelaphus buxtoni

Tragelaphus spekii

Tragelaphus scriptus

Tragelaphus imberbis

Tragelaphus eurycerus

Bison bison

Bos mutus

Bison bonasus

Bos primigenius

Bos sauveli

Bos javanicus

Bos gaurus

Pseudoryx nghetinhensis

Syncerus caffer

Pelorovis antiquus

Bubalus mindorensis

Bubalus palaeokerabau

Bubalus arnee

Bubalus depressicornis

Bubalus quarlesi

Aepyceros melampus

Nesotragus moschatus

Neotragus batesi

Neotragus pygmaeus

Philantomba maxwellii

Philantomba monticola

Sylvicapra grimmia

Cephalophus zebra

Cephalophus jentinki

Cephalophus dorsalis

Cephalophus silvicultor

Cephalophus spadix

Cephalophus adersi

Cephalophus niger

Cephalophus ogilbyi

Cephalophus weynsi

Cephalophus callipygus

Cephalophus leucogaster

Cephalophus harveyi

Cephalophus natalensis

Cephalophus rufilatus

Cephalophus nigrifrons

Raphicerus campestris

Raphicerus melanotis

Raphicerus sharpei

Dorcatragus megalotis

Madoqua guentheri

Madoqua kirkii

Madoqua saltiana

Madoqua piacentinii

Ourebia ourebi

Procapra picticaudata

Procapra gutturosa

Procapra przewalskii

Saiga tatarica

Litocranius walleri

Antidorcas bondi

Antidorcas marsupialis

Antidorcas australis

Ammodorcas clarkei

Antilope cervicapra

Nanger dama

Nanger soemmerringii

Nanger granti

Spirocerus kiakhtensis

Eudorcas thomsonii

Eudorcas albonotata

Eudorcas rufifrons

Eudorcas rufina

Gazella gazella

Gazella spekei

Gazella saudiya

Gazella dorcas

Gazella bilkis

Gazella subgutturosa

Gazella bennettii

Gazella leptoceros

Gazella tingitana

Gazella cuvieri

Gazella atlantica

Oreotragus oreotragus

Pelea capreolus

Redunca fulvorufula

Redunca redunca

Redunca arundinum

Kobus ellipsiprymnus

Kobus megaceros

Kobus leche

Kobus kob

Kobus vardonii

Hippotragus niger

Hippotragus equinus

Hippotragus leucophaeus

Addax nasomaculatus

Oryx beisa

Oryx gazella

Oryx dammah

Oryx leucoryx

Rusingoryx atopocranion

Connochaetes taurinus

Connochaetes gnou

Beatragus hunteri

Alcelaphus buselaphus

Megalotragus priscus

Damaliscus lunatus

Damaliscus pygargus

Damaliscus hypsodon

Damaliscus niro

Pantholops hodgsonii

Capricornis crispus

Ovibos moschatus

Capricornis sumatraensis

Capricornis swinhoei

Capricornis milneedwardsii

Capricornis thar

Capricornis rubidus

Megalovis guangxiensis

Bootherium bombifrons

Euceratherium collinum

Naemorhedus goral

Naemorhedus caudatus

Naemorhedus griseus

Naemorhedus baileyi

Oreamnos americanus

Oreamnos harringtoni

Rupicapra rupicapra

Rupicapra pyrenaica

Myotragus balearicus

Budorcas taxicolor

Nilgiritragus hylocrius

Ovis ammon

Ovis orientalis

Ovis nivicola

Ovis canadensisOvis dalli

Arabitragus jayakari

Ammotragus lervia

Caprini indet

Pseudois schaeferi

Pseudois nayaur

Hemitragus jemlahicus

Hemitragus cedrensis

Soergelia minor

Capra sibirica

Capra ibex

Capra pyrenaica

Capra walie

Capra nubiana

Capra falconeri

Capra aegagrus

Capra caucasica

Capra cylindricornis

Rhinopoma microphyllum

Rhinopoma muscatellum

Rhinopoma hardwickii

Rhinopoma macinnesi

Craseonycteris thonglongyai

Cardioderma cor

Macroderma gigas

Lavia frons

Megaderma lyra

Megaderma spasma

Rhinolophus rufus

Rhinolophus landeri

Rhinolophus guineensis

Rhinolophus alcyone

Rhinolophus yunanensis

Rhinolophus blasii

Rhinolophus mehelyi

Rhinolophus euryale

Rhinolophus denti

Rhinolophus simulator

Rhinolophus swinnyi

Rhinolophus capensis

Rhinolophus virgo

Rhinolophus euryotis

Rhinolophus sedulus

Rhinolophus ferrumequinum

Rhinolophus celebensis

Rhinolophus madurensis

Rhinolophus paradoxolophus

Rhinolophus clivosus

Rhinolophus bocharicus

Rhinolophus hillorum

Rhinolophus ruwenzorii

Rhinolophus maclaudi

Rhinolophus hilli

Rhinolophus ziama

Rhinolophus darlingi

Rhinolophus hildebrandti

Rhinolophus inops

Rhinolophus trifoliatus

Rhinolophus fumigatus

Rhinolophus eloquens

Rhinolophus maendeleo

Rhinolophus hipposideros

Rhinolophus mitratus

Rhinolophus formosae

Rhinolophus beddomei

Rhinolophus luctus

Rhinolophus pearsonii

Rhinolophus cognatus

Rhinolophus subbadius

Rhinolophus affinis

Rhinolophus creaghi

Rhinolophus canuti

Rhinolophus osgoodi

Rhinolophus stheno

Rhinolophus shameli

Rhinolophus subrufus

Rhinolophus arcuatus

Rhinolophus deckenii

Rhinolophus thomasi

Rhinolophus sinicus

Rhinolophus rouxii

Rhinolophus malayanus

Rhinolophus acuminatus

Rhinolophus borneensis

Rhinolophus nereis

Rhinolophus megaphyllus

Rhinolophus adami

Rhinolophus pusillus

Rhinolophus shortridgei

Rhinolophus lepidus

Rhinolophus sakejiensis

Rhinolophus rex

Rhinolophus philippinensis

Rhinolophus montanus

Rhinolophus marshalli

Rhinolophus coelophyllus

Rhinolophus silvestris

Rhinolophus keyensis

Rhinolophus robinsoni

Rhinolophus macrotis

Rhinolophus siamensis

Rhinolophus convexus

Asellia tridens

Asellia patrizii

Aselliscus stoliczkanus

Aselliscus tricuspidatus

Paracoelops megalotis

Coelops hirsutus

Coelops frithii

Coelops robinsoni

Cloeotis percivali

Rhinonicteris aurantia

Triaenops auritus

Triaenops furculus

Triaenops persicus

Triaenops rufus

Hipposideros gigas

Hipposideros commersoni

Hipposideros thomensis

Hipposideros vittatus

Hipposideros camerunensis

Hipposideros cyclops

Hipposideros corynophyllus

Hipposideros stenotis

Hipposideros wollastoni

Hipposideros semoni

Hipposideros muscinus

Hipposideros edwardshilli

Hipposideros boeadii

Hipposideros speoris

Anthops ornatus

Hipposideros jonesi

Hipposideros obscurus

Hipposideros megalotis

Hipposideros cervinus

Hipposideros galeritus

Hipposideros coxi

Hipposideros pygmaeus

Hipposideros pratti

Hipposideros scutinares

Hipposideros lylei

Hipposideros armiger

Hipposideros turpis

Hipposideros sorenseni

Hipposideros madurae

Hipposideros grandis

Hipposideros sumbae

Hipposideros larvatus

Hipposideros lekaguli

Hipposideros inexpectatus

Hipposideros dinops

Hipposideros pelingensis

Hipposideros lankadiva

Hipposideros inornatus

Hipposideros diadema

Hipposideros demissus

Hipposideros calcaratus

Hipposideros coronatus

Hipposideros beatus

Hipposideros curtus

Hipposideros macrobullatus

Hipposideros ruber

Hipposideros lamottei

Hipposideros abae

Hipposideros fuliginosus

Hipposideros papua

Hipposideros caffer

Hipposideros breviceps

Hipposideros fulvus

Hipposideros ridleyi

Hipposideros marisae

Hipposideros ater

Hipposideros dyacorum

Hipposideros crumeniferus

Hipposideros doriae

Hipposideros halophyllus

Hipposideros orbiculus

Hipposideros khaokhouayensis

Hipposideros rotalis

Hipposideros pomona

Hipposideros nequam

Hipposideros cineraceus

Hipposideros durgadasi

Hipposideros maggietaylorae

Hipposideros bicolor

Hipposideros hypophyllus

Eidolon helvum

Eidolon dupreanum

Pteropus leucopterus

Pteropus insularis

Pteropus pselaphon

Pteropus subniger

Notopteris macdonaldi

Notopteris neocaledonica

Paranyctimene raptor

Paranyctimene tenax

Nyctimene aello

Nyctimene certans

Nyctimene cyclotis

Nyctimene draconilla

Nyctimene minutus

Nyctimene albiventer

Nyctimene keasti

Nyctimene robinsoni

Nyctimene major

Nyctimene cephalotes

Nyctimene malaitensis

Nyctimene masalai

Nyctimene rabori

Nyctimene vizcaccia

Nyctimene sanctacrucis

Melonycteris melanops

Melonycteris woodfordi

Melonycteris fardoulisi

Pteralopex taki

Pteralopex anceps

Pteralopex flanneryi

Pteralopex atrata

Pteralopex pulchra

Mirimiri acrodonta

Neopteryx frosti

Styloctenium wallacei

Styloctenium mindorensis

Acerodon mackloti

Acerodon leucotis

Acerodon celebensis

Acerodon humilis

Acerodon jubatus

Pteropus molossinus

Pteropus woodfordi

Pteropus mahaganus

Pteropus tokudae

Pteropus neohibernicus

Pteropus tonganus

Pteropus pelewensis

Pteropus yapensis

Pteropus ualanus

Pteropus lombocensis

Pteropus mariannus

Pteropus speciosus

Pteropus hypomelanus

Pteropus admiralitatum

Pteropus conspicillatus

Pteropus ocularis

Pteropus alecto

Pteropus griseus

Pteropus pohlei

Pteropus macrotis

Pteropus nitendiensis

Pteropus vetulus

Pteropus tuberculatus

Pteropus temminckii

Pteropus anetianus

Pteropus cognatus

Pteropus chrysoproctus

Pteropus rayneri

Pteropus rennelli

Pteropus fundatus

Pteropus pilosus

Pteropus samoensis

Pteropus brunneus

Pteropus capistratus

Pteropus poliocephalus

Pteropus ornatus

Pteropus voeltzkowi

Pteropus personatus

Pteropus caniceps

Pteropus howensis

Pteropus livingstonii

Pteropus aruensis

Pteropus keyensis

Pteropus melanopogon

Pteropus coxi

Pteropus scapulatus

Pteropus gilliardorum

Pteropus dasymallus

Pteropus pumilus

Pteropus faunulus

Pteropus rodricensis

Pteropus vampyrus

Pteropus intermedius

Pteropus giganteus

Pteropus lylei

Pteropus argentatus

Pteropus allenorum

Pteropus melanotus

Pteropus niger

Pteropus seychellensis

Pteropus aldabrensis

Pteropus rufus

Pteropus loochoensis

Dyacopterus spadiceus

Dyacopterus brooksi

Sphaerias blanfordi

Balionycteris maculata

Aethalops alecto

Aethalops aequalis

Latidens salimalii

Penthetor lucasi

Chironax melanocephalus

Thoopterus nigrescens

Alionycteris paucidentata

Otopteropus cartilagonodus

Haplonycteris fischeri

Ptenochirus jagori

Ptenochirus minor

Megaerops wetmorei

Megaerops ecaudatus

Megaerops niphanae

Megaerops kusnotoi

Cynopterus nusatenggara

Cynopterus horsfieldii

Cynopterus titthaecheilus

Cynopterus sphinx

Cynopterus brachyotis

Cynopterus luzoniensis

Cynopterus minutus

Macroglossus minimus

Macroglossus sobrinus

Syconycteris hobbit

Syconycteris australis

Syconycteris carolinae

Rousettus bidens

Aproteles bulmerae

Harpyionycteris celebensis

Harpyionycteris whiteheadi

Dobsonia emersa

Dobsonia minor

Dobsonia peronii

Dobsonia moluccensis

Dobsonia anderseni

Dobsonia exoleta

Dobsonia chapmani

Dobsonia pannietensis

Dobsonia praedatrix

Dobsonia inermis

Dobsonia beauforti

Dobsonia viridis

Dobsonia crenulata

Casinycteris argynnis

Scotonycteris zenkeri

Scotonycteris ophiodon

Rousettus aegyptiacus

Rousettus spinalatus

Rousettus amplexicaudatus

Rousettus linduensis

Rousettus leschenaultii

Rousettus madagascariensis

Rousettus obliviosus

Rousettus celebensis

Eonycteris spelaea

Eonycteris robusta

Eonycteris major

Rousettus lanosus

Megaloglossus woermanni

Lissonycteris angolensis

Myonycteris relicta

Myonycteris brachycephala

Myonycteris torquata

Plerotes anchietae

Hypsignathus monstrosus

Epomops dobsonii

Epomops franqueti

Epomops buettikoferi

Nanonycteris veldkampii

Micropteropus pusillus

Micropteropus intermedius

Epomophorus wahlbergi

Epomophorus angolensis

Epomophorus grandis

Epomophorus anselli

Epomophorus labiatus

Epomophorus minimus

Epomophorus gambianus

Epomophorus crypturus

Mystacina tuberculata

Mystacina robusta

Thyroptera discifera

Thyroptera lavali

Thyroptera tricolor

Thyroptera devivoi

Amorphochilus schnablii

Furipterus horrens

Noctilio leporinus

Noctilio albiventris

Mormoops megalophylla

Mormoops blainvillei

Mormoops magna

Pteronotus parnellii

Pteronotus paraguanensis

Pteronotus personatus

Pteronotus macleayii

Pteronotus quadridens

Pteronotus gymnonotus

Pteronotus davyi

Lampronycteris brachyotis

Micronycteris schmidtorum

Micronycteris minuta

Micronycteris hirsuta

Micronycteris sanborni

Micronycteris brosseti

Micronycteris megalotis

Micronycteris microtis

Micronycteris matses

Macrotus californicus

Macrotus waterhousii

Diphylla ecaudata

Diaemus youngi

Desmodus stocki

Desmodus puntajudensis

Desmodus rotundus

Desmodus draculae

Vampyrum spectrum

Chrotopterus auritus

Trachops cirrhosus

Macrophyllum macrophyllum

Phyllostomus discolorTonatia bidens

Tonatia saurophila

Lophostoma carrikeri

Lophostoma schulzi

Lophostoma brasiliense

Lophostoma evotis

Lophostoma silvicolum

Lophostoma yasuni

Lophostoma aequatorialis

Phylloderma stenops

Phyllostomus latifolius

Phyllostomus hastatus

Phyllostomus elongatus

Mimon cozumelaeMimon bennettii

Mimon crenulatum

Mimon koepckeae

Lonchorhina orinocensis

Lonchorhina fernandezi

Lonchorhina inusitata

Lonchorhina aurita

Lonchorhina marinkellei

Anoura fistulata

Anoura luismanueli

Anoura caudiferAnoura cultrata

Anoura latidens

Anoura geoffroyi

Hylonycteris underwoodi

Musonycteris harrisoni

Lichonycteris obscura

Scleronycteris ega

Choeronycteris mexicana

Choeroniscus minor

Choeroniscus godmani

Choeroniscus periosus

Erophylla bombifrons

Erophylla sezekorni

Brachyphylla cavernarum

Brachyphylla nana

Phyllonycteris aphylla

Phyllonycteris poeyi

Phyllonycteris major

Monophyllus redmani

Monophyllus plethodon

Leptonycteris nivalis

Leptonycteris curasoae

Leptonycteris yerbabuenae

Glossophaga commissarisi

Glossophaga soricina

Glossophaga leachii

Glossophaga longirostris

Glossophaga morenoi

Lonchophylla orcesi

Lonchophylla mordax

Lonchophylla thomasi

Lonchophylla handleyi

Lonchophylla bokermanni

Lonchophylla chocoana

Lionycteris spurrelli

Platalina genovensium

Xeronycteris vieirai

Lonchophylla concava

Lonchophylla robusta

Lonchophylla hesperia

Lonchophylla dekeyseri

Trinycteris nicefori

Neonycteris pusilla

Glyphonycteris daviesi

Glyphonycteris sylvestris

Glyphonycteris behnii

Carollia perspicillata

Carollia castanea

Carollia brevicaudaCarollia subrufaCarollia sowelliCarollia manu

Rhinophylla fischerae

Rhinophylla pumilio

Rhinophylla alethinaSturnira bidens

Sturnira sorianoiSturnira nana

Sturnira aratathomasi

Sturnira erythromosSturnira luisi

Sturnira magna

Sturnira oporaphilumSturnira ludovici

Sturnira thomasiSturnira mordax

Sturnira bogotensisSturnira lilium

Sturnira tildae

Sturnira mistratensis

Vampyressa bidens

Vampyressa nymphaea

Vampyressa brocki

Chiroderma salvini

Chiroderma doriae

Chiroderma trinitatum

Chiroderma villosum

Chiroderma improvisum

Uroderma magnirostrum

Uroderma bilobatum

Mesophylla macconnelli

Vampyressa pusilla

Vampyressa thyone

Vampyressa melissa

Platyrrhinus lineatus

Platyrrhinus recifinus

Platyrrhinus chocoensis

Platyrrhinus helleri

Platyrrhinus matapalensis

Platyrrhinus brachycephalus

Vampyrodes caraccioli

Platyrrhinus vittatus

Platyrrhinus albericoi

Platyrrhinus aurarius

Platyrrhinus nigellusPlatyrrhinus masu

Platyrrhinus ismaeli

Platyrrhinus dorsalis

Platyrrhinus infuscus

Platyrrhinus umbratus

Enchisthenes hartiiEctophylla alba

Stenoderma rufumPhyllops falcatus

Phyllops silvai

Ariteus flavescensArdops nichollsi

Sphaeronycteris toxophyllumCenturio senex

Pygoderma bilabiatumAmetrida centurio

Artibeus phaeotisArtibeus toltecus

Artibeus aztecusArtibeus cinereus

Artibeus anderseniArtibeus gnomus

Artibeus glaucus

Artibeus rosenbergiiArtibeus concolorArtibeus watsoni

Artibeus incomitatusArtibeus anthonyi

Artibeus inopinatusArtibeus hirsutus

Artibeus fraterculus

Artibeus jamaicensisArtibeus lituratus

Artibeus fimbriatusArtibeus obscurus

Artibeus planirostrisArtibeus amplus

Nycteris thebaica

Nycteris gambiensis

Nycteris madagascariensisNycteris macrotisNycteris vinsoniNycteris woodi

Nycteris parisiiNycteris grandis

Nycteris hispidaNycteris aurita

Nycteris javanicaNycteris tragataNycteris nanaNycteris arge

Nycteris majorNycteris intermedia

Saccolaimus peli

Saccolaimus mixtus

Saccolaimus saccolaimus

Saccolaimus flaviventris

Taphozous hamiltoni

Taphozous nudiventris

Taphozous hildegardeae

Taphozous perforatus

Taphozous longimanus

Taphozous mauritianus

Taphozous melanopogonTaphozous achates

Taphozous theobaldiTaphozous hilli

Taphozous kapalgensisTaphozous australis

Taphozous georgianus

Taphozous troughtoniMosia nigrescens

Emballonura atrataEmballonura tiavato

Coleura afra

Coleura seychellensisEmballonura serii

Emballonura beccarii

Emballonura semicaudata

Emballonura raffrayanaEmballonura dianaeEmballonura furax

Emballonura alecto

Emballonura monticola

Rhynchonycteris naso

Centronycteris centralis

Centronycteris maximiliani

Saccopteryx canescens

Saccopteryx gymnuraSaccopteryx leptura

Saccopteryx bilineata

Saccopteryx antioquensisBalantiopteryx plicata

Balantiopteryx infuscaBalantiopteryx io

Peropteryx leucopteraPeropteryx trinitatis

Peropteryx kappleriPeropteryx macrotis

Cormura brevirostrisCyttarops alecto

Diclidurus isabellusDiclidurus scutatusDiclidurus ingensDiclidurus albus

Myzopoda aurita

Myzopoda schliemanni

Chilonatalus tumidifronsChilonatalus micropus

Nyctiellus lepidusNatalus stramineus

Natalus lanatusNatalus tumidirostrisNatalus mexicanus

Natalus primusNatalus jamaicensis

Natalus espiritosantensisNatalus major

Tomopeas ravusPlatymops setiger

Mormopterus beccarii

Mormopterus norfolkensis

Mormopterus planicepsMormopterus loriae

Mormopterus doriaeMormopterus jugularis

Mormopterus acetabulosusMormopterus phrudus

Mormopterus kalinowskiiMormopterus minutus

Cheiromeles torquatusCheiromeles parvidensMyopterus daubentonii

Myopterus whitleyi

Molossops aequatorianus

Molossops mattogrossensisMolossops neglectus

Molossops temminckiiCynomops abrasus

Cynomops paranusCynomops planirostrisCynomops greenhalli

Cynomops mexicanusOtomops martiensseni

Otomops madagascariensisOtomops formosus

Otomops johnstoneiOtomops papuensis

Otomops wroughtoniOtomops secundus

Sauromys petrophilusNyctinomops macrotis

Nyctinomops aurispinosus

Nyctinomops femorosaccus

Nyctinomops laticaudatusPromops centralisPromops nasutus

Molossus sinaloaeMolossus currentium

Molossus rufusMolossus pretiosus

Molossus molossusMolossus barnesi

Molossus coibensisMolossus aztecus

Eumops underwoodiEumops auripendulusEumops bonariensis

Eumops patagonicusEumops hansaeEumops perotis

Eumops trumbulliEumops dabbeneiEumops maurus

Eumops glaucinusEumops floridanusTadarida australis

Tadarida kuboriensisTadarida fulminans

Tadarida lobataTadarida ventralis

Tadarida latoucheiTadarida teniotis

Tadarida insignisTadarida brasiliensisTadarida aegyptiacaTadarida jobimenaTadarida russata

Tadarida gallagheriTadarida tomensisTadarida nigeriaeTadarida plicataTadarida major

Tadarida midasTadarida mops

Tadarida niangaraeTadarida congicaTadarida pumila

Tadarida aloysiisabaudiaeTadarida bivittata

Tadarida ansorgeiTadarida bemmeleni

Tadarida jobensisTadarida chapini

Tadarida solomonisTadarida johorensisTadarida bregullae

Tadarida trevoriTadarida sarasinorumTadarida leucostigma

Tadarida demonstratorTadarida niveiventerTadarida condyluraTadarida petersoniTadarida thersitesTadarida nanula

Tadarida brachypteraTadarida spurrelli

Miniopterus fuliginosusMiniopterus schreibersii

Miniopterus tristisMiniopterus robustior

Miniopterus magnaterMiniopterus shortridgei

Miniopterus oceanensisMiniopterus macrocneme

Miniopterus pusillusMiniopterus paululus

Miniopterus natalensisMiniopterus sororculus

Miniopterus newtoniMiniopterus inflatusMiniopterus minor

Miniopterus fraterculusMiniopterus fuscus

Miniopterus mediusMiniopterus griveaudi

Miniopterus petersoniMiniopterus manavi

Miniopterus gleniMiniopterus majori

Miniopterus australisEudiscopus denticulus

Cistugo seabraeCistugo lesueuri

Rhogeessa gracilisRhogeessa io

Rhogeessa hussoniRhogeessa genowaysi

Rhogeessa alleniScotophilus kuhlii

Scotophilus collinusScotophilus nux

Scotophilus marovazaScotophilus heathii

Scotophilus celebensisScotophilus leucogaster

Scotophilus nucellaScotophilus tandrefana

Scotophilus robustusScotophilus nigritaScotophilus viridis

Scotophilus borbonicusScotophilus dinganii

Harpiola isodonHarpiola grisea

Harpiocephalus harpiaHarpiocephalus mordax

Murina putaMurina huttoni

Murina leucogasterMurina aurata

Murina hilgendorfiMurina fusca

Murina cyclotisMurina tubinaris

Murina aeneaMurina rozendaali

Murina floriumMurina suilla

Murina tenebrosaMurina ussuriensis

Murina harrisoniMurina ryukyuana

Phoniscus papuensisPhoniscus atrox

Phoniscus jagoriiKerivoula agnellaKerivoula lanosa

Kerivoula cuprosaKerivoula myrella

Kerivoula lenisKerivoula papillosa

Kerivoula kachinensisKerivoula flora

Kerivoula minutaKerivoula argentata

Kerivoula titaniaKerivoula intermediaKerivoula eriophoraKerivoula phalaenaKerivoula muscina

Kerivoula krauensisKerivoula pellucida

Kerivoula whiteheadiKerivoula africana

Kerivoula hardwickiiKerivoula smithiiKerivoula picta

Myotis alcathoeMyotis hajastanicus

Myotis brandtiiMyotis insularum

Myotis volansMyotis sodalis

Myotis fortidensMyotis vivesi

Myotis auriculusMyotis septentrionalis

Myotis csorbaiMyotis californicus

Myotis leibiiMyotis planiceps

Myotis ciliolabrumMyotis melanorhinus

Myotis aelleniMyotis lucifugusMyotis occultus

Myotis keeniiMyotis evotis

Myotis thysanodesMyotis keaysiMyotis ruber

Myotis simusMyotis riparius

Myotis elegansMyotis austroriparius

Myotis grisescensMyotis velifer

Myotis yumanensisMyotis cobanensisMyotis chiloensis

Myotis oxyotusMyotis levis

Myotis findleyiMyotis nigricans

Myotis nesopolusMyotis peninsularis

Myotis albescensMyotis pilosus

Myotis martiniquensisMyotis atacamensis

Myotis dominicensisMyotis moluccarum

Myotis tricolorMyotis emarginatus

Myotis morrisiMyotis scotti

Myotis bocagiiMyotis welwitschiiMyotis formosusMyotis hermani

Myotis anjouanensisMyotis goudoti

Myotis rufopictusMyotis dinellii

Myotis dasycnemeMyotis muricola

Myotis aterMyotis annectans

Myotis gomantongensisMyotis pruinosus

Myotis montivagusMyotis yanbarensis

Myotis nipalensisMyotis mystacinusMyotis aurascensMyotis ikonnikovi

Myotis dieteriMyotis altarium

Myotis oreiasMyotis sicarius

Myotis bechsteiniiMyotis daubentonii

Myotis fraterMyotis bucharensis

Myotis punicusMyotis myotisMyotis blythii

Myotis chinensisMyotis schaubi

Myotis pequiniusMyotis nattereri

Myotis bombinusMyotis adversus

Myotis fimbriatusMyotis davidii

Myotis siligorensisMyotis laniger

Myotis longipesMyotis annamiticus

Myotis capacciniiMyotis rosseti

Myotis australisMyotis ridleyi

Myotis macropusMyotis hasseltii

Myotis horsfieldiiMyotis macrodactylus

Myotis macrotarsusMyotis stalkeri

Idionycteris phyllotisNycticeius humeralisNycticeius cubanus

Euderma maculatumLasiurus xanthinus

Lasiurus egaLasiurus intermedius

Lasiurus insularisLasiurus ebenus

Lasiurus egregiusLasiurus cinereus

Lasiurus castaneusLasiurus atratus

Lasiurus degelidusLasiurus seminolus

Lasiurus minorLasiurus blossevillii

Lasiurus variusLasiurus borealisLasiurus pfeifferi

Barbastella barbastellusBarbastella leucomelas

Antrozous pallidusBauerus dubiaquercus

Rhogeessa parvulaRhogeessa mira

Rhogeessa minutillaRhogeessa tumida

Rhogeessa aeneusOtonycteris hemprichii

Corynorhinus rafinesquiiCorynorhinus townsendiiCorynorhinus mexicanus

Plecotus auritusPlecotus ognevi

Plecotus macrobullarisPlecotus sardus

Plecotus taivanusPlecotus sacrimontis

Plecotus christiiPlecotus balensis

Plecotus austriacusPlecotus kolombatovici

Plecotus teneriffaePipistrellus hesperus

Pipistrellus musciculusPipistrellus subflavus

Pipistrellus joffreiPipistrellus anthonyi

Pipistrellus stenopterusLasionycteris noctivagans

Arielulus aureocollarisArielulus cuprosusArielulus societatis

Arielulus circumdatusArielulus torquatus

Glauconycteris alboguttataGlauconycteris variegata

Glauconycteris machadoiGlauconycteris kenyacola

Glauconycteris egeriaGlauconycteris curryae

Glauconycteris superbaGlauconycteris poensisGlauconycteris beatrix

Glauconycteris humeralisGlauconycteris gleni

Glauconycteris argentataScotomanes ornatus

Ia ioScotozous dormeri

Scoteanax rueppelliiFalsistrellus mackenziei

Falsistrellus tasmaniensisFalsistrellus mordax

Falsistrellus affinisFalsistrellus petersiEptesicus nasutusEptesicus matroka

Eptesicus malagasyensisEptesicus hottentotus

Eptesicus serotinusEptesicus bottae

Eptesicus bobrinskoiEptesicus nilssonii

Eptesicus gobiensisEptesicus japonensis

Histiotus macrotusHistiotus laephotis

Histiotus velatusHistiotus humboldti

Histiotus alienusHistiotus magellanicus

Histiotus montanusEptesicus floweriEptesicus fuscus

Eptesicus pachyotisEptesicus chiriquinus

Eptesicus andinusEptesicus brasiliensis

Eptesicus furinalisEptesicus diminutusEptesicus innoxiusEptesicus platyops

Eptesicus tateiEptesicus dimissus

Eptesicus guadeloupensisHesperoptenus tickelli

Hesperoptenus tomesiHesperoptenus gaskelliHesperoptenus doriae

Hesperoptenus blanfordiPhiletor brachypterus

Tylonycteris pachypusTylonycteris robustulaLaephotis botswanae

Laephotis namibensisLaephotis wintoni

Laephotis angolensisPipistrellus capensis

Pipistrellus somalicusPipistrellus zuluensis

Pipistrellus imbricatusPipistrellus arabicus

Pipistrellus heliosPipistrellus nanus

Pipistrellus rendalliPipistrellus flavescensPipistrellus tenuipinnisPipistrellus melckorum

Pipistrellus brunneusPipistrellus guineensisPipistrellus eisentrautiPipistrellus anchietae

Pipistrellus pulveratusPipistrellus kitcheneriPipistrellus lophurus

Nycticeinops schlieffeniScotorepens balstoni

Scotorepens orionScotorepens greyii

Scotorepens sanborniPipistrellus cadornaePipistrellus macrotis

Pipistrellus vordermanniPipistrellus ariel

Pipistrellus alaschanicusPipistrellus savii

Chalinolobus tuberculatusChalinolobus gouldii

Chalinolobus neocaledonicusChalinolobus nigrogriseus

Chalinolobus dwyeriChalinolobus morio

Chalinolobus picatusVespadelus caurinus

Vespadelus vulturnusVespadelus finlaysoni

Vespadelus baverstockiVespadelus pumilus

Vespadelus troughtoniVespadelus douglasorum

Vespadelus darlingtoniVespadelus regulusMimetillus moloneyiVespertilio murinusVespertilio sinensisScotoecus pallidusScotoecus hirundo

Scotoecus albofuscusGlischropus tylopus

Glischropus javanusPipistrellus nathusii

Nyctalus plancyi

Nyctalus furvus

Nyctalus leisleri

Nyctalus azoreum

Nyctalus noctula

Nyctalus montanus

Nyctalus lasiopterus

Nyctalus aviator

Pharotis imogene

Nyctophilus heran

Nyctophilus nebulosus

Nyctophilus walkeri

Nyctophilus microtis

Nyctophilus microdon

Nyctophilus timoriensis

Nyctophilus sherrini

Nyctophilus geoffroyi

Nyctophilus arnhemensis

Nyctophilus howensis

Nyctophilus gouldi

Nyctophilus bifax Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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collin

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Pipist

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Euoti

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Galag

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Galag

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Galag

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Galag

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Eulem

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Eulem

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Eulem

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Eulem

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Eulem

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Micro

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Micro

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Micro

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Micro

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Tarsi

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Tarsi

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Tarsi

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Caca

jao m

elano

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alus

Caca

jao ay

resi

Chiro

potes

chiro

potes

Chiro

potes

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asCh

iropo

tes ut

ahick

aeCh

iropo

tes al

binas

usAn

tillothr

ix be

rnens

isXe

nothr

ix mc

grego

riCa

lliceb

us ni

grifro

nsCa

lliceb

us co

imbra

iCa

lliceb

us ba

rbarab

rowna

eCa

lliceb

us pe

rsona

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lliceb

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elano

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Callic

ebus

molo

chCa

lliceb

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neras

cens

Callic

ebus

brun

neus

Callic

ebus

hoffm

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iCa

lliceb

us ba

ptista

Callic

ebus

bernh

ardi

Callic

ebus

dubiu

sCa

lliceb

us au

reipa

latii

Callic

ebus

cupre

usCa

lliceb

us di

scolo

rCa

lliceb

us or

natus

Callic

ebus

step

henn

ashi

Callic

ebus

calig

atus

Callic

ebus

oena

nthe

Callic

ebus

mod

estus

Callic

ebus

olall

aeCa

lliceb

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naco

philu

sCa

lliceb

us pa

llesc

ens

Callic

ebus

lucif

erCa

lliceb

us m

edem

iCa

lliceb

us pu

rinus

Callic

ebus

luge

nsCa

lliceb

us re

gulus

Callic

ebus

torqu

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Aloua

tta pi

graAlo

uatta

pallia

taAlo

uatta

guari

baAlo

uatta

nige

rrima

Aloua

tta be

lzebu

lAlo

uatta

ulula

taAlo

uatta

disc

olor

Aloua

tta ca

raya

Aloua

tta se

niculu

sAlo

uatta

juara

Aloua

tta pu

ruens

isAlo

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sara

Aloua

tta ar

ctoide

aAlo

uatta

mac

conn

elli

Ateles

panis

cus

Ateles

cham

ekAte

les m

argina

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les fu

scice

psAte

les hy

bridu

sAte

les ge

offroy

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les be

lzebu

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achy

teles

arac

hnoid

esBr

achy

teles

hypo

xanth

usOr

eona

x flav

icaud

aCa

ipora

bamb

uiorum

Proto

pithe

cus b

rasilie

nsis

Lago

thrix

lugen

sLa

gothr

ixlag

otrich

aLa

gothr

ix ca

naLa

gothr

ix po

eppig

iiSa

imiri

bolivi

ensis

Saim

iri va

nzoli

niiSa

imiri

ustus

Saim

iri sc

iureu

sSa

imiri

oerst

edii

Cebu

s oliva

ceus

Cebu

s kaa

pori

Cebu

s albi

frons

Cebu

s cap

ucinu

sCe

bus a

pella

Cebu

s xan

thoste

rnos

Cebu

s rob

ustus

Cebu

s flav

iusCe

bus n

igritu

sCe

bus m

acroc

epha

lusCe

bus l

ibidin

osus

Cebu

s cay

Aotus

nigri

ceps

Aotus

jorge

herna

ndez

iAo

tus m

icona

xAo

tus vo

cifera

nsAo

tus zo

nalis

Aotus

trivir

gatus

Aotus

grise

imem

braAo

tus le

murin

usAo

tus na

ncym

aae

Aotus

azara

eAo

tus br

umba

cki

Sagu

inus l

euco

pus

Sagu

inus t

ripart

itus

Sagu

inus n

igrico

llisSa

guinu

s mela

noleu

cus

Sagu

inus f

uscic

ollis

Sagu

inus i

nustu

sSa

guinu

s imp

erator

Sagu

inus l

abiat

usSa

guinu

s mys

taxSa

guinu

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ffroyi

Sagu

inus o

edipu

sSa

guinu

s bico

lorSa

guinu

s mart

insi

Sagu

inus n

iger

Sagu

inus m

idas

Leon

topith

ecus

chrys

omela

sLe

ontop

ithec

us ro

salia

Leon

topith

ecus

chrys

opyg

usLe

ontop

ithec

us ca

issara

Callim

ico go

eldii

Callib

ella h

umilis

Callith

rix au

ritaCa

llithrix

flavic

eps

Callith

rix ge

offroy

iCa

llithrix

kuhli

iCa

llithrix

jacc

hus

Callith

rix pe

nicilla

taCe

buell

a pyg

maea

Mico

sater

eiMi

co ch

rysole

ucus

Mico

mau

esi

Mico

hume

ralife

rMi

co le

ucipp

eMi

co m

arcai

Mico

acari

ensis

Mico

man

icoren

sisMi

co ni

grice

psMi

co ar

genta

tusMi

co in

terme

dius

Mico

mela

nurus

Mico

emilia

eMi

co ro

ndon

iPo

ngo a

belii

Pong

o pyg

maeu

sGo

rilla b

ering

eiGo

rilla g

orilla

Pan p

anisc

usPa

n trog

lodyte

sHo

mo er

ectus

Homo

flores

iensis

Homo

Den

isova

nsHo

mo sa

piens

Homo

nean

derth

alens

isSy

mpha

langu

s syn

dacty

lusNo

masc

us na

sutus

Noma

scus

haina

nus

Noma

scus

conc

olor

Noma

scus

gabri

ellae

Noma

scus

leuc

ogen

ysNo

masc

us si

kiHo

olock

leuc

oned

ysHo

olock

hoolo

ckHy

lobate

s pile

atus

Hylob

ates l

arHy

lobate

s molo

chHy

lobate

s klos

siiHy

lobate

s mue

lleri

Hylob

ates a

lbiba

rbis

Hylob

ates a

gilis

Colob

us sa

tanas

Colob

us an

golen

sisCo

lobus

guere

zaCo

lobus

velle

rosus

Colob

us po

lykom

osPr

ocolo

bus v

erus

Proc

olobu

s rufo

mitra

tusPr

ocolo

bus k

irkii

Proc

olobu

s gord

onoru

mPr

ocolo

bus b

adius

Proc

olobu

s pen

nanti

iPr

ocolo

bus p

reuss

iPr

esby

tis th

omas

iPr

esby

tis po

tenzia

niPr

esby

tis co

mata

Pres

bytis

rubic

unda

Pres

bytis

mela

lopho

sPr

esby

tis fro

ntata

Pres

bytis

hose

iPr

esby

tis ch

rysom

elas

Pres

bytis

femo

ralis

Pres

bytis

siam

ensis

Pres

bytis

natun

aeRh

inopit

hecu

s brel

ichi

Rhino

pithe

cus r

oxell

ana

Rhino

pithe

cus a

vunc

ulus

Rhino

pithe

cus b

ieti

Nasa

lis lar

vatus

Simias

conc

olor

Pyga

thrix

nigrip

esPy

gathr

ix ne

maeu

sPy

gathr

ix cin

erea

Trach

ypith

ecus

hatin

hens

isTra

chyp

ithec

us la

otum

Trach

ypith

ecus

delac

ouri

Trach

ypith

ecus

polio

ceph

alus

Trach

ypith

ecus

franc

oisi

Trach

ypith

ecus

phay

reiTra

chyp

ithec

us ob

scuru

sTra

chyp

ithec

us ba

rbei

Trach

ypith

ecus

crist

atus

Trach

ypith

ecus

aurat

usTra

chyp

ithec

us ge

rmain

iTra

chyp

ithec

us pi

leatus

Trach

ypith

ecus

short

ridge

iTra

chyp

ithec

us ge

eiTra

chyp

ithec

us jo

hnii

Trach

ypith

ecus

vetul

usSe

mnop

ithec

us du

ssum

ieri

Semn

opith

ecus

entel

lusSe

mnop

ithec

us hy

poleu

cos

Semn

opith

ecus

schis

taceu

s

Semn

opith

ecus

hecto

rSe

mnop

ithec

us aj

axSe

mnop

ithec

us pr

iamAll

enop

ithec

us ni

grovir

idis

Eryth

roceb

us pa

tasCe

rcopit

hecu

s sola

tusCe

rcopit

hecu

s lho

esti

Cerco

pithe

cus p

reuss

iCh

loroc

ebus

cyno

suros

Chlor

oceb

us py

geryt

hrus

Chlor

oceb

us ta

ntalus

Chlor

oceb

us ae

thiop

sCh

loroc

ebus

saba

eus

Chlor

oceb

us dj

amdja

mens

is

Miop

ithec

us og

ouen

sisMi

opith

ecus

talap

oinCe

rcopit

hecu

s ham

lyni

Cerco

pithe

cus n

eglec

tusCe

rcopit

hecu

s pog

onias

Cerco

pithe

cus c

ampb

elli

Cerco

pithe

cus m

ona

Cerco

pithe

cus d

iana

Cerco

pithe

cus d

ryas

Cerco

pithe

cus m

itisCe

rcopit

hecu

s nict

itans

Cerco

pithe

cus a

scan

iusCe

rcopit

hecu

s cep

hus

Cerco

pithe

cus s

clater

iCe

rcopit

hecu

s peta

urista

Cerco

pithe

cus e

rythro

gaste

r

Cerco

pithe

cus e

rythro

tisMa

ndrill

us sp

hinx

Mand

rillus

leuc

opha

eus

Cerco

cebu

s torq

uatus

Cerco

cebu

s san

jeiCe

rcoce

bus a

tysCe

rcoce

bus c

hryso

gaste

r

Cerco

cebu

s agil

isCe

rcoce

bus g

aleritu

sTh

eropit

hecu

s gela

daLo

phoc

ebus

aterr

imus

Loph

oceb

us al

bigen

aRu

ngwe

cebu

s kipu

njiPa

pio pa

pioPa

pio ha

madry

asPa

pio cy

noce

phalu

sPa

pio ur

sinus

Papio

anub

isMa

caca

sylva

nus

Maca

ca fa

scicu

laris

Maca

ca m

ulatta

Maca

ca cy

clopis

Maca

ca fu

scata

Maca

ca ar

ctoide

sMa

caca

thibe

tana

Maca

ca as

same

nsis

Maca

ca m

unza

laMa

caca

sinic

aMa

caca

radia

taMa

caca

page

nsis

Maca

ca si

beru

Maca

ca si

lenus

Maca

ca le

onina

Maca

ca ne

mestr

inaMa

caca

nigre

scen

sMa

caca

heck

iMa

caca

mau

raMa

caca

tonk

eana

Maca

ca ni

graMa

caca

ochre

ataPti

locerc

us lo

wiiDe

ndrog

ale m

elanu

raDe

ndrog

ale m

urina

Anath

ana e

llioti

Urog

ale ev

eretti

Tupa

ia gli

sTu

paia

belan

geri

Tupa

ia lon

gipes

Tupa

ia ch

rysog

aster

Tupa

ia gra

cilis

Tupa

ia do

rsalis

Tupa

ia jav

anica

Tupa

ia nic

obari

caTu

paia

palaw

anen

sisTu

paia

moell

endo

rffiTu

paia

mino

rTu

paia

tana

Tupa

ia pic

taTu

paia

splen

didula

Tupa

ia mo

ntana

Prolag

us sa

rdus

Ocho

tona m

uliens

isOc

hoton

a glov

eriOc

hoton

a eryt

hrotis

Ocho

tona f

orres

tiOc

hoton

a gao

ligon

gens

is

Ocho

tona r

ufesce

nsOc

hoton

a mac

rotis

Ocho

tona r

oylei

Ocho

tona h

imala

yana

Ocho

tona k

oslow

iOc

hoton

a ruti

laOc

hoton

a ilien

sisOc

hoton

a lad

acen

sisOc

hoton

a nigr

itiaOc

hoton

a hua

ngen

sisOc

hoton

a pus

illaOc

hoton

a tho

masi

Ocho

tona d

auuri

caOc

hoton

a thib

etana

Ocho

tona c

ansu

sOc

hoton

a curz

oniae

Ocho

tona n

ubric

aOc

hoton

a colla

risOc

hoton

a prin

ceps

Ocho

tona w

harto

niOc

hoton

a turu

chan

ensis

Ocho

tona a

lpina

Ocho

tona a

rgenta

taOc

hoton

a palla

siOc

hoton

a hoff

mann

iOc

hoton

a hyp

erbore

a

Capro

lagus

hisp

idus

Oryct

olagu

s cun

iculus

Buno

lagus

mon

ticular

is

Penta

lagus

furne

ssiBra

chyla

gus id

ahoe

nsis

Sylvil

agus

palus

trisSy

lvilag

us in

sonu

sSy

lvilag

us aq

uaticu

sSy

lvilag

us m

ansu

etus

Sylvil

agus

bach

mani

Sylvil

agus

bras

iliens

is

Sylvil

agus

dice

iSy

lvilag

us va

rynae

nsis

Sylvil

agus

audu

bonii

Sylvil

agus

nutta

lliiSy

lvilag

us tra

nsitio

nalis

Sylvil

agus

obscu

rusSy

lvilag

us cu

nicula

rius

Sylvil

agus

gray

soni

Sylvil

agus

robu

stus

Sylvil

agus

florid

anus

Sylvil

agus

cogn

atus

Rome

rolag

us di

azi

Neso

lagus

netsc

heri

Neso

lagus

timmi

nsi

Poela

gus m

arjori

taPro

nolag

us cr

assic

auda

tus

Prono

lagus

rand

ensis

Prono

lagus

rupe

stris

Prono

lagus

saun

dersi

ae

Aztla

nolag

us ag

ilisLe

pus s

inens

isLe

pus m

ands

huric

us

Lepu

s gran

atens

isLe

pus e

uropa

eus

Lepu

s hain

anus

Lepu

s nigr

icollis

Lepu

s peg

uens

isLe

pus o

iostol

usLe

pus c

omus

Lepu

s ame

rican

usLe

pus b

rachy

urus

Lepu

s fag

ani

Lepu

s sax

atilis

Lepu

s micr

otis

Lepu

s yark

ande

nsis

Lepu

s cors

icanu

sLe

pus c

apen

sisLe

pus h

abes

sinicu

s

Lepu

s star

ckiLe

pus t

olai

Lepu

s tibe

tanus

Lepu

s callo

tisLe

pus f

lavigu

laris

Lepu

s allen

iLe

pus c

aliforn

icus

Lepu

s insu

laris

Lepu

s tow

nsen

diiLe

pus c

astro

viejoi

Lepu

s othu

sLe

pus a

rcticu

sLe

pus c

orean

usLe

pus t

imidu

sLa

onas

tes ae

nigma

mus

Felov

ia va

eMa

ssouti

era m

zabi

Pectin

ator s

peke

iCte

noda

ctylus

gund

i

Cteno

dacty

lus va

liTri

chys

fascic

ulata

Atheru

rus m

acrou

rus

Atheru

rus af

rican

us

Hystr

ix cris

tata

Hystr

ix afric

aeau

stralis

Hystr

ix ind

icaHy

strix b

rachy

uraHy

strix j

avan

icaHy

strix c

rassis

pinis

Hystr

ix pum

ilaHy

strix r

efossa

Hystr

ix kian

gsen

ensis

Hystr

ix sum

atrae

Petro

mus t

ypicu

sTh

ryono

mys s

winde

rianu

s

Thryo

nomy

s greg

orian

us

Heter

ocep

halus

glab

er

Heliop

hobiu

s arge

nteoc

inereu

s

Geory

chus

cape

nsis

Bathy

ergus

suillu

sBa

thyerg

us ja

netta

Cryp

tomys

hotte

ntotus

Cryp

tomys

foxi

Cryp

tomys

ochra

ceoc

inereu

s

Cryp

tomys

mech

owi

Cryp

tomys

zech

iCr

yptom

ys bo

cage

i

Cryp

tomys

darlin

gi

Cryp

tomys

dama

rensis

Cryp

tomys

anse

lli

Cryp

tomys

kafue

nsis

Chae

tomys

subs

pinos

us

Erethi

zon d

orsatu

m

Echin

oproc

ta ruf

esce

ns

Coen

dou b

icolor

Coen

dou n

ycthe

mera

Coen

dou p

rehen

silis

Coen

dou q

uichu

aCo

endo

u san

ctama

rtae

Coen

dou r

othsch

ildi

Sphig

gurus

mex

icanu

s

Sphig

gurus

spino

sus

Sphig

gurus

villos

us

Sphig

gurus

vestit

us

Sphig

gurus

pruin

osus

Sphig

gurus

mela

nurus

Sphig

gurus

ichillu

s

Sphig

gurus

insid

iosus

Sphig

gurus

roos

malen

orum

Cunic

ulus t

acza

nows

kii

Cunic

ulus p

aca

Myop

rocta

acou

chy

Myop

rocta

pratti

Dasyp

rocta

kalino

wskii

Dasyp

rocta

ruatan

ica

Dasyp

rocta

lepori

na

Dasyp

rocta

prymn

oloph

a

Dasyp

rocta

fuligin

osa

Dasyp

rocta

azara

e

Dasyp

rocta

coiba

e

Dasyp

rocta

guam

ara

Dasyp

rocta

punc

tata

Dasyp

rocta

mexic

ana

Dasyp

rocta

crista

ta

Dolich

otis p

atago

num

Dolich

otis s

alinico

la

Kerod

on ru

pestr

is

Kerod

on ac

robata

Neoc

hoeru

s aes

opi

Hydro

choe

rus hy

droch

aeris

Hydro

choe

rus ist

hmius

Galea

mon

aster

iensis

Galea

mus

teloid

es

Galea

spixii

Galea

flavid

ens

Micro

cavia

austr

alis

Micro

cavia

niata

Micro

cavia

shipt

oni

Cavia

mag

naCa

via fu

lgida

Cavia

apere

aCa

via ts

chud

iiCa

via in

terme

dia

Dinom

ys bra

nickii

Lago

stomu

s max

imus

Lago

stomu

s cras

sus

Chinc

hilla c

hinch

illa

Chinc

hilla l

anige

ra

Lagid

ium pe

ruanu

m

Lagid

ium vis

cacia

Lagid

ium wo

lffsoh

ni

Cusco

mys a

shan

inka

Cusco

mys o

blativa

Abroc

oma b

enne

ttii

Abroc

oma b

olivien

sis

Abroc

oma c

inerea

Abroc

oma s

histac

ea

Abroc

oma v

acca

rum

Abroc

oma b

udini

Abroc

oma f

amati

na

Abroc

oma u

spalla

ta

Tymp

anoc

tomys

barre

rae

Pipan

acoc

tomys

aureu

s

Octom

ys mi

max

Octod

ontom

ys glir

oides

Spala

copu

s cya

nus

Acon

aemy

s port

eri

Acon

aemy

s sag

ei

Acon

aemy

s fus

cus

Salino

ctomy

s losch

alcha

leros

orum

Octod

on lu

natus

Octod

on br

idges

i

Octod

on de

gus

Octod

on pa

cificu

s

Cteno

mys f

lamari

oni

Cteno

mys a

ustra

lis

Cteno

mys p

orteo

usi

Cteno

mys a

zarae

Cteno

mys b

ergi

Cteno

mys y

oland

ae

Cteno

mys b

onett

oi

Cteno

mys r

ioneg

rensis

Cteno

mys d

orsalis

Cteno

mys m

endo

cinus

Cteno

mys f

ochi

Cteno

mys p

undti

Cteno

mys t

alarum

Cteno

mys b

rasilie

nsis

Cteno

mys t

orqua

tus

Cteno

mys la

miCte

nomy

s minu

tus

Cteno

mys p

earso

ni

Cteno

mys r

oigi

Cteno

mys d

orbign

yi

Cteno

mys p

erren

si

Cteno

mys o

svaldo

reigi

Cteno

mys m

agella

nicus

Cteno

mys h

aigi

Cteno

mys c

oyha

iquen

sis

Cteno

mys f

odax

Cteno

mys s

erice

us

Cteno

mys t

ucum

anus

Cteno

mys p

eruan

us

Cteno

mys ju

risCte

nomy

s latro

Cteno

mys p

ilaren

sis

Cteno

mys a

rgenti

nus

Cteno

mys o

ccultu

s

Cteno

mys m

aulinu

s

Cteno

mys s

teinb

achi

Cteno

mys b

olivien

sis

Cteno

mys g

oodfe

llowi

Cteno

mys le

ucod

on

Cteno

mys t

ucon

ax

Cteno

mys s

ociab

ilis

Cteno

mys c

onov

eri

Cteno

mys le

wisi

Cteno

mys e

milian

us

Cteno

mys v

alidus

Cteno

mys f

rater

Cteno

mys b

udini

Cteno

mys s

ylvan

us

Cteno

mys s

caglia

i

Cteno

mys s

altari

us

Cteno

mys c

olburn

i

Cteno

mys p

ontife

x

Cteno

mys o

pimus

Cteno

mys k

nighti

Cteno

mys v

iperin

us

Cteno

mys tu

lduco

Cteno

mys fa

mosu

s

Cteno

mys jo

hann

is

Cteno

mys c

oludo

Cteno

mys fu

lvus

Hexo

lobod

on ph

enax

Hexo

lobod

ontina

eGen

NOW

Sp NO

W

Ambly

rhiza

inund

ata

Quem

isia gr

avis

Xaym

aca fu

lvopu

lvis

Isolob

odon

mon

tanus

Isolob

odon

porto

ricens

is

Rhizo

plagio

dontia

lemkei

Plagio

donti

a aed

ium

Plagio

dontia

ipnae

um

Capro

mys p

iloride

s

Capro

mys S

P Now

Geoca

promy

s brow

nii

Geoca

promy

s thora

catus

Geoca

promy

s SP A

Geoca

promy

s SP B

Geoca

promy

s ingra

hami

Geoca

promy

s colu

mbian

us

Mesoc

aprom

ys na

nus

Mesoc

aprom

ys au

ritus

Mesoc

aprom

ys san

felipe

nsis

Mesoc

aprom

ys an

gelca

brerai

Mysat

eles g

arrido

i

Mysat

eles m

eridio

nalis

Mysat

eles m

elanu

rus

Mysat

eles g

undla

chi

Mysat

eles p

rehen

silis

Myoca

stor c

oypus

Hoplo

mys g

ymnu

rus

Proech

imys p

oliopu

s

Proech

imys c

hrysae

olus

Proech

imys d

ecuma

nus

Proech

imys o

conne

lli

Proech

imys lo

ngica

udatu

s

Proech

imys e

chino

thrix

Proech

imys s

emisp

inosus

Proech

imys b

revica

uda

Proech

imys g

oeldii

Proech

imys q

uadru

plicatu

s

Proech

imys c

uvieri

Proech

imys r

obert

i

Proech

imys p

atton

i

Proech

imys tr

initatu

s

Proech

imys k

ulinae

Proech

imys c

anico

llis

Proech

imys g

uyann

ensis

Proech

imys s

teerei

Proech

imys h

oplom

yoide

s

Proech

imys s

imonsi

Proech

imys g

ardne

ri

Proech

imys u

richi

Proech

imys m

incae

Proech

imys g

uaira

e

Proech

imys m

agda

lenae

Thrich

omys

inerm

is

Thrich

omys

apere

oides

Thrich

omys

pachy

urus

Euryz

ygoma

tomys

spino

sus

Carte

rodon

sulcid

ens

Clyom

ys lati

ceps

Clyom

ys bis

hopi

Trino

mys g

ratios

us

Trino

mys ih

ering

i

Trino

mys d

imidia

tus

Trino

mys m

yosuro

s

Trino

mys y

onen

agae

Trino

mys a

lbispin

us

Trino

mys s

etosus

Trino

mys e

liasi

Trino

mys m

oojen

i

Trino

mys p

aratus

Trino

mys m

irapita

nga

Lonch

othrix

emilia

e

Isothr

ix pag

urus

Isothr

ix sinn

amari

ensis

Isothr

ix neg

rensis

Isothr

ix bistr

iata

Mesom

ys his

pidus

Mesom

ys occ

ultus

Mesom

ys len

iceps

Mesom

ys stim

ulax

Brotom

ys vor

atus

Heter

opsom

ys ins

ulans

Borom

ys off

ella

Borom

ys tor

rei

Kann

abate

omys

ambly

onyx

Olallam

ys alb

icaud

a

Olallam

ys ed

ax

Dacty

lomys

dacty

linus

Dacty

lomys

perua

nus

Dacty

lomys

bolivi

ensis

Santa

marta

mys r

ufodo

rsalis

Diplom

ys can

iceps

Diplom

ys lab

ilis

Callis

tomys

pictus

Makal

ata ob

scura

Makal

ata did

elpho

ides

Makal

ata m

acrura

Makal

ata rh

ipidura

Torom

ys gra

ndis

Patto

nomy

s occa

sius

Patto

nomy

s sem

ivillos

us

Echim

ys chr

ysurus

Echim

ys sat

urnus

Echim

ys vie

irai

Phyllo

mys n

igrisp

inus

Phyllo

mys d

asythr

ix

Phyllo

mys u

nicolo

r

Phyllo

mys th

omasi

Phyllo

mys m

antiqu

eiren

sis

Phyllo

mys p

atton

i

Phyllo

mys lu

ndi

Phyllo

mys la

marum

Phyllo

mys k

erri

Phyllo

mys b

rasilie

nsis

Phyllo

mys b

lainvill

ii

Phyllo

mys m

edius

Musca

rdinu

s avel

lanari

us

Selev

inia be

tpakda

laensi

s

Myom

imus r

oachi

Myom

imus p

erson

atus

Myom

imus s

etzeri

Chae

tocau

da sic

huan

ensis

Dryom

ys lan

iger

Dryom

ys nite

dula

Dryom

ys nie

thamm

eri

Eliom

ys qu

ercinu

s

Eliom

ys mo

rpheu

s

Eliom

ys me

lanuru

s

Eliom

ys mu

nbyan

us

Glis g

lisGli

rulus

japon

icus

Grap

hiurus

nagtg

lasii

Grap

hiurus

crass

icaud

atus

Grap

hiurus

platyo

ps

Grap

hiurus

ango

lensis

Grap

hiurus

mon

ardi

Grap

hiurus

ocula

ris

Grap

hiurus

christ

yi

Grap

hiurus

micro

tis

Grap

hiurus

muri

nus

Grap

hiurus

lorrai

neus

Grap

hiurus

kellen

i

Grap

hiurus

johnst

oni

Grap

hiurus

rupic

ola

Grap

hiurus

surdu

s

Aplod

ontia

rufa

Sperm

ophilo

psis le

ptoda

ctylus

Xerus

rutilu

s

Xerus

eryth

ropus

Atlan

toxeru

s getu

lus

Xerus

princ

eps

Xerus

inauri

s

Helios

ciurus

ruwe

nzorii

Helios

ciurus

undu

latus

Helios

ciurus

gamb

ianus

Helios

ciurus

punct

atus

Helios

ciurus

rufob

rachiu

m

Helios

ciurus

muta

bilis

Epixe

rus eb

ii

Protox

erus s

tange

ri

Protox

erus a

ubinn

ii

Myosc

iurus

pumil

io

Funis

ciurus

carru

thersi

Funis

ciurus

lemnis

catus

Funis

ciurus

subst

riatus

Funis

ciurus

pyrro

pus

Funis

ciurus

leucog

enys

Funis

ciurus

cong

icus

Funis

ciurus

bayon

ii

Funis

ciurus

anery

thrus

Funis

ciurus

isabe

lla

Funis

ciurus

ducha

illui

Parax

erus c

oope

ri

Parax

erus p

alliatu

s

Parax

erus b

oehm

i

Parax

erus v

incen

ti

Parax

erus v

exillar

ius

Parax

erus o

chrace

us

Parax

erus p

oensi

s

Parax

erus c

epap

i

Parax

erus a

lexan

dri

Parax

erus fl

avovitt

is

Parax

erus lu

cifer

Sciur

otamia

s davi

dianu

s

Sciur

otamia

s forre

sti

Tamia

s sibir

icus

Tamia

s minim

us

Tamia

s spe

ciosus

Tamia

s qua

drima

culatu

s

Tamia

s pana

mintinu

s

Tamias

alpinu

s

Tamias

duran

gae

Tamias

buller

i

Tamias

canip

es

Tamias

rufus

Tamias

quadr

ivittatu

s

Tamias

dorsa

lis

Tamias

cinere

icollis

Tamias

palmeri

Tamias

umbrin

us

Tamias

merriam

i

Tamias

obscu

rus

Tamias

amoen

us

Tamias

rufica

udus

Tamias

siskiy

ou

Tamias

senex

Tamias

sonom

ae

Tamias

ochro

genys

Tamias

striatu

s

Tamias

townse

ndii

Ammosp

ermoph

ilus int

erpres

Ammosp

ermoph

ilus ha

rrisii

Ammosp

ermoph

ilus leu

curus

Ammosp

ermoph

ilus ne

lsoni

Sperm

ophilus

varieg

atus

Sperm

ophilus

beech

eyi

Sperm

ophilus

atrica

pillus

Sperm

ophilus

latera

lis

Sperm

ophilus

satur

atus

Sperm

ophilus

madrens

is

Marmota

flaviv

entris

Marmota

olympus

Marmota

vanco

uveren

sis

Marmota

caliga

ta

Marmota

monax

Marmota

brow

eri

Marmota

marmota

Marmota

menzbie

ri

Marmota

cauda

ta

Marmota

bobak

Marmota

baiba

cina

Marmota

camtsc

hatica

Marmota

himala

yana

Marmota

sibiric

a

Cynom

ys mexi

canus

Cynom

ys lud

ovicia

nus

Cynom

ys par

videns

Cynom

ys leu

curus

Cynom

ys gun

nisoni

Sperm

ophilus

annul

atus

Sperm

ophilus

adoce

tus

Sperm

ophilus

musicus

Sperm

ophilus

pygm

aeus

Sperm

ophilus

major

Sperm

ophilus

dauric

us

Sperm

ophilus

suslic

us

Sperm

ophilus

xanth

oprym

nus

Sperm

ophilus

citellu

s

Sperm

ophilus

tauren

sis

Sperm

ophilus

eryth

rogeny

s

Sperm

ophilus

fulvus

Sperm

ophilus

brevi

cauda

Sperm

ophilus

relict

us

Sperm

ophilus

ralli

Sperm

ophilus

pallid

icauda

Sperm

ophilus

alasha

nicus

Sperm

ophilus

mexican

us

Sperm

ophilus

tridece

mlineatu

s

Sperm

ophilus

frankl

inii

Sperm

ophilus

perot

ensis

Sperm

ophilus

spilos

oma

Sperm

ophilus

teretic

audus

Sperm

ophilus

mohaven

sis

Sperm

ophilus

beldin

gi

Sperm

ophilus

washi

ngtoni

Sperm

ophilus

townse

ndii

Sperm

ophilus

brunn

eus

Sperm

ophilus

canus

Sperm

ophilus

mollis

Sperm

ophilus

armatu

s

Sperm

ophilus

colum

bianus

Sperm

ophilus

undul

atus

Sperm

ophilus

parry

ii

Sperm

ophilus

richar

dsonii

Sperm

ophilus

elegan

s

Funam

bulus

tristria

tus

Funam

bulus

pennan

tii

Funam

bulus

palmaru

m

Funam

bulus

subline

atus

Funam

bulus

layard

i

Exilis

ciurus

exilis

Exilis

ciurus

conci

nnus

Exilis

ciurus

whiteh

eadi

Hyosc

iurus h

einrich

i

Hyosc

iurus il

eile

Rubris

ciurus

rubriv

enter

Prosci

urillus

abstru

sus

Prosci

urillus

leucom

us

Prosci

urillus

rosen

bergii

Prosci

urillus

murinus

Prosci

urillus

weber

i

Rhino

sciuru

s latica

udatus

Tamiop

s rodol

phii

Tamiop

s macc

lelland

ii

Tamiop

s swin

hoei

Tamiop

s marit

imus

Dremom

ys lok

riah

Dremom

ys per

nyi

Dremom

ys gul

aris

Dremom

ys ruf

igenis

Dremom

ys pyr

rhomeru

s

Dremom

ys eve

retti

Sunda

sciuru

s frate

rculus

Sunda

sciuru

s lowii

Sunda

sciuru

s broo

kei

Sunda

sciuru

s tenui

s

Sunda

sciuru

s jentin

ki

Sunda

sciuru

s stee

rii

Sunda

sciuru

s juven

cus

Sunda

sciuru

s dave

nsis

Sunda

sciuru

s rabor

i

Sunda

sciuru

s sam

arensi

s

Sunda

sciuru

s moel

lendor

ffi

Sunda

sciuru

s hipp

urus

Sunda

sciuru

s mind

anensi

s

Sunda

sciuru

s hoog

straali

Sunda

sciuru

s phili

ppinen

sis

Nanno

sciuru

s mela

notis

Callos

ciurus

prevo

stii

Callos

ciurus

pyger

ythrus

Callos

ciurus

nigrov

ittatus

Callos

ciurus

albesc

ens

Menetes

berdm

orei

Larisc

us ins

ignis

Larisc

us hos

ei

Larisc

us obs

curus

Larisc

us nio

be

Glyphot

es sim

us

Callos

ciurus

quinq

uestria

tus

Callos

ciurus

melanog

aster

Callos

ciurus

phayr

ei

Callos

ciurus

balue

nsis

Callos

ciurus

inorna

tus

Callos

ciurus

notatu

s

Callos

ciurus

adam

si

Callos

ciurus

oreste

s

Callos

ciurus

finlay

sonii

Callos

ciurus

canic

eps

Callos

ciurus

erythra

eus

Sciuril

lus pu

sillus

Ratufa

macrour

a

Ratufa

indica

Ratufa

affinis

Ratufa

bicolo

r

Tamias

ciurus

hudso

nicus

Tamias

ciurus

mearnsi

Tamias

ciurus

dougl

asii

Rheith

rosciu

rus macr

otis

Synthe

osciuru

s broc

hus

Sciuru

s stram

ineus

Sciuru

s ignitu

s

Microsci

urus a

lfari

Microsci

urus s

antand

erensi

s

Microsci

urus fl

aviven

ter

Microsci

urus m

imulus

Sciuru

s aber

ti

Sciuru

s gilvig

ularis

Sciuru

s anom

alus

Sciuru

s varie

gatoid

es

Sciuru

s flam

mifer

Sciuru

s gran

atensi

s

Sciuru

s caro

linensi

s

Sciuru

s nige

r

Sciuru

s depp

ei

Sciuru

s naya

ritensi

s

Sciuru

s puch

eranii

Sciuru

s yuca

tanens

is

Sciuru

s vulg

aris

Sciuru

s aest

uans

Sciuru

s pyrrh

inus

Sciuru

s aure

ogaste

r

Sciuru

s lis

Sciuru

s sanb

orni

Sciuru

s colli

aei

Sciuru

s richm

ondi

Sciuru

s allen

i

Sciuru

s grise

us

Sciuru

s ocul

atus

Sciuru

s arizo

nensis

Sciuru

s igniv

entris

Sciuru

s spad

iceus

Petau

rista le

ucogen

ys

Petau

rista x

anthot

is

Petau

rista p

hilippe

nsis

Petau

rista e

legans

Petau

rista p

etauris

ta

Petau

rista m

agnific

us

Petau

rista a

lborufu

s

Petau

rista n

obilis

Pterom

ys vol

ans

Pterom

ys mom

onga

Eupet

aurus

cinere

us

Aerom

ys tep

hromela

s

Aerom

ys tho

masi

Pterom

yscus

pulver

ulentu

s

Aerete

s mela

nopteru

s

Trogop

terus x

anthip

es

Belom

ys pea

rsonii

Biswam

oyopte

rus bis

wasi

Eogla

ucomys

fimbria

tus

Glaucom

ys vol

ans

Glaucom

ys sab

rinus

Iomys

horsfie

ldii

Iomys

sipora

Petino

mys gen

ibarbis

Petino

mys hag

eni

Petino

mys crin

itus

Petino

mys mind

anensi

s

Petino

mys sag

itta

Petino

mys vor

dermann

i

Petino

mys lug

ens

Petino

mys set

osus

Petino

mys fus

cocapi

llus

Petau

rillus h

osei

Petau

rillus k

inlochi

i

Petau

rillus e

miliae

Hylopet

es pha

yrei

Hylopet

es wins

toni

Hylopet

es alb

oniger

Hylopet

es nig

ripes

Hylopet

es spa

diceus

Hylopet

es sip

ora

Hylopet

es bar

telsi

Hylopet

es lep

idus

Hylopet

es pla

tyurus

Castoro

ides o

hioens

is

Castor c

anaden

sis

Castor f

iber

Thomom

ys mont

icola

Thomom

ys clu

sius

Thomom

ys talp

oides

Thomom

ys maza

ma

Thomom

ys ida

hoensi

s

Thomom

ys bul

bivoru

s

Thomom

ys tow

nsendi

i

Thomom

ys bot

tae

Thomom

ys um

brinus

Geomys

attwate

ri

Geomys

tropica

lis

Geomys

person

atus

Geomys

pinetis

Geomys

brevic

eps

Geomys

bursar

ius

Geomys

texens

is

Geomys

knoxjo

nesi

Geomys

arenar

ius

Zygoge

omys

trichop

us

Orthogeo

mys his

pidus

Orthogeo

mys lan

ius

Orthogeo

mys und

erwood

i

Orthogeo

mys cav

ator

Orthogeo

mys che

rriei

Orthogeo

mys mata

galpae

Orthogeo

mysdar

iensis

Orthogeo

mys het

erodus

Orthogeo

mys tha

eleri

Orthogeo

mys gra

ndis

Orthogeo

mys cun

iculus

Pappog

eomys

bulleri

Pappog

eomys

alcorn

i

Cratogeo

mys fum

osus

Cratogeo

mys pla

niceps

Cratogeo

mys cas

tanops

Cratogeo

mys gol

dmani

Cratogeo

mys per

otensi

s

Cratogeo

mys fulv

escens

Cratogeo

mys merri

ami

Microdip

odops

megacep

halus

Microdip

odops

pallidu

s

Dipodom

ys des

erti

Dipodom

ys ord

ii

Dipodom

ys spe

ctabilis

Dipodom

ys nel

soni

Dipodom

ys com

pactus

Dipodom

ys phi

llipsii

Dipodom

ys ela

tor

Dipodom

ys nitr

atoide

s

Dipodom

ys merri

ami

Dipodom

ys ins

ularis

Dipodom

ys marg

aritae

Dipodom

ys cal

ifornic

us

Dipodom

ys ven

ustus

Dipodom

ys sim

ulans

Dipodom

ys agi

lis

Dipodom

ys ing

ens

Dipodom

ys gra

vipes

Dipodom

ys micro

ps

Dipodom

ys ste

phensi

Dipodom

ys hee

rmanni

Dipodom

ys pan

aminti

nus

Liomys

salvin

i

Liomys

adsper

sus

Liomys

irroratu

s

Liomys

pictus

Liomys

specta

bilis

Heterom

ys nel

soni

Heterom

ys ore

sterus

Heterom

ys gau

meri

Heterom

ys oas

icus

Heterom

ys aus

tralis

Heterom

ys ano

malus

Heterom

ys des

marestia

nus

Heterom

ys tele

us

Perogna

thus fa

sciatus

Perogna

thus fl

avesce

ns

Perogna

thus a

lticolus

Perogna

thus p

arvus

Perogna

thus fl

avus

Perogna

thus m

erriam

i

Perogna

thus a

mplus

Perogna

thus in

ornatu

s

Perogna

thus lo

ngimem

bris

Chaetod

ipus ru

dinoris

Chaetod

ipus b

aileyi

Chaetod

ipus fo

rmosus

Chaetod

ipus h

ispidu

s

Chaetod

ipus in

termedi

us

Chaetod

ipus n

elsoni

Chaetod

ipus lin

eatus

Chaetod

ipus g

oldmani

Chaetod

ipus a

rtus

Chaetod

ipus p

enicilla

tus

Chaetod

ipus e

remicu

s

Chaetod

ipus p

ernix

Chaetod

ipus s

pinatu

s

Chaetod

ipus c

aliforn

icus

Chaetod

ipus fa

llax

Chaetod

ipus d

alques

ti

Chaetod

ipus a

renariu

s

Pedetes

capen

sis

Pedetes

surda

ster

Zenker

ella ins

ignis

Idiurus

macrotis

Idiurus

zenke

ri

Anomalu

rus de

rbianus

Anomalu

rus pu

sillus

Anomalu

rus be

ecrofti

Anomalu

rus pe

lii

Sicista s

trandi

Sicista c

audata

Sicista a

rmenica

Sicista c

oncolo

r

Sicista c

aucasi

ca

Sicista s

evertzo

vi

Sicista t

iansha

nica

Sicista n

apaea

Sicista k

azbegi

ca

Sicista b

etulina

Sicista p

seudon

apaea

Sicista s

ubtilis

Sicista k

luchoric

a

Allactod

ipus b

obrinsk

ii

Pygeretm

us pla

tyurus

Pygeretm

us pum

ilio

Pygeretm

us zhit

kovi

Euchore

utes n

aso

Eozapus

setch

uanus

Napaeoz

apus in

signis

Zapus

hudson

ius

Zapus

trinota

tus

Zapus

princep

s

Allactag

a euph

ratica

Allactag

a willia

msi

Allactag

a sibir

ica

Allactag

a balik

unica

Allactag

a bulla

ta

Allactag

a tetrad

actyla

Allactag

a elate

r

Allactag

a firou

zi

Allactag

a seve

rtzovi

Allactag

a hots

oni

Allactag

a majo

r

Allactag

a vino

gradov

i

Paradip

us cte

nodact

ylus

Cardiocr

anius

parado

xus

Salping

otulus

michaelis

Salping

otus ko

zlovi

Salping

otus h

eptner

i

Salping

otus p

allidus

Salping

otus cr

assicau

da

Eremodi

pus lich

tenste

ini

Dipus sa

gitta

Jaculu

s orien

talis

Jaculu

s jaculu

s

Jaculu

s blan

fordi

Stylodip

us sun

gorus

Stylodip

us and

rewsi

Stylodip

us telu

m

Platacan

thomys l

asiurus

Typhlo

mys cine

reus

Eospala

x roths

childi

Eospala

x fonta

nierii

Eospala

x smithii

Myospal

ax psil

urus

Myospal

ax asp

alax

Myospal

ax myos

palax

Tachyo

ryctes

splen

dens

Tachyo

ryctes

macroce

phalus

Rhizomys s

inensis

Rhizomys s

umatre

nsis

Rhizomys p

ruinosu

s

Cannom

ys badi

us

Spalax

ehrenb

ergi

Spalax

nehring

i

Spalax

leucod

on

Spalax

graecu

s

Spalax

micropht

halmus

Spalax

zemni

Spalax

arenar

ius

Spalax

gigant

eus

Spalax

uralen

sis

Calomysc

us hot

soni

Calomysc

us mysta

x

Calomysc

us elb

urzens

is

Calomysc

us bai

lwardi

Calomysc

us gra

ndis

Calomysc

us tso

lovi

Calomysc

us ura

rtensis

Calomysc

us bal

uchi

Monticol

omys k

oopmani

Macrota

rsomys p

etteri

Macrota

rsomys b

astard

i

Macrota

rsomys i

ngens

Hypogeo

mys antim

ena

Hypogeo

mys aust

ralis

Brachyu

romys r

amiroh

itra

Brachyu

romys b

etsileo

ensis

Nesomys a

udeber

ti

Nesomys r

ufus

Nesomys l

amber

toni

Brachyt

arsom

ys albic

auda

Brachyt

arsom

ys villos

a

Gymnurom

ys robe

rti

Voalavo g

ymnocaud

us

Voalavo a

ntsaha

bensis

Eliurus gr

andidie

ri

Eliurus an

tsingy

Eliurus m

ajori

Eliurus da

nieli

Eliurus ta

nala

Eliurus w

ebbi

Eliurus el

lermani

Eliurus m

inor

Eliurus pe

tteri

Eliurus m

yoxinus

Eliurus pe

nicillatu

s

Mystrom

ys albic

audatu

s

Delanymys b

rooksi

Petromysc

us collin

us

Petromysc

us barb

ouri

Petromysc

us mont

icularis

Petromysc

us shor

tridgei

Beamys h

indei

Cricetom

ys gam

bianus

Cricetom

ys emini

Saccosto

mus cam

pestris

Saccosto

mus mear

nsi

Malacoth

rix typic

a

Prionom

ys bate

si

Dendrop

rionom

ys rous

seloti

Steatom

ys kreb

sii

Steatom

ys cupp

edius

Steatom

ys jack

soni

Steatom

ys caur

inus

Steatom

ys prate

nsis

Steatom

ys parv

us

Steatom

ys boca

gei

Steatom

ys opim

us

Megaden

dromus n

ikolaus

i

Dendrom

us kivu

Dendrom

us orea

s

Dendrom

us vern

ayi

Dendrom

us kahu

ziensis

Dendrom

us mysta

calis

Dendrom

us insig

nis

Dendrom

us mess

orius

Dendrom

us nyika

e

Dendrom

us meso

melas

Dendrom

us melan

otis

Dendrom

us lova

ti

Nelsonia

goldmani

Nelsonia

neotom

odon

Hodomys a

lleni

Xenomys n

elsoni

Neotom

a ciner

ea

Neotom

a mexic

ana

Neotom

a palat

ina

Neotom

a goldm

ani

Neotom

a albig

ula

Neotom

a steph

ensi

Neotom

a chryso

melas

Neotom

a brya

nti

Neotom

a lepida

Neotom

a devia

Neotom

a phen

ax

Neotom

a fuscip

es

Neotom

a macr

otis

Neotom

a magis

ter

Neotom

a marti

nensis

Neotom

a florida

na

Neotom

a nelso

ni

Neotom

a micro

pus

Neotom

a angu

stapalat

a

Neotom

a leuco

don

Neotom

a bunk

eri

Neotoma an

thonyi

Ochrotom

ys nutta

lli

Scotinom

ys xera

mpelinus

Scotinom

ys tegu

ina

Baiomys m

usculus

Baiomys t

aylori

Isthmomys p

irrensis

Isthmomys f

lavidus

Reithrod

ontomys c

hrysops

is

Reithrod

ontomys m

ontanus

Reithrod

ontomys r

aviventr

is

Reithrod

ontomys h

umulis

Reithrod

ontomys h

irsutus

Reithrod

ontomys f

ulvesce

ns

Reithrod

ontomys s

umichrasti

Reithrod

ontomys z

acateca

e

Reithrod

ontomys m

egalotis

Reithrod

ontomys b

urti

Reithrod

ontomys c

reper

Reithrod

ontomys d

arienen

sis

Reithrod

ontomys m

exicanu

s

Reithrod

ontomys b

revirost

ris

Reithrod

ontomys s

pectabi

lis

Reithrod

ontomys g

racilis

Reithrod

ontomys p

aradox

us

Reithrod

ontomys t

enuiros

tris

Reithrod

ontomys b

akeri

Reithrod

ontomys m

icrodon

Reithrod

ontomys r

odrigue

zi

Onychom

ys aren

icola

Onychom

ys leuc

ogaster

Onychom

ys torri

dus

Megadon

tomys c

ryophilu

s

Megadon

tomys t

homasi

Megadon

tomys n

elsoni

Osgoodo

mys band

eranus

Peromysc

us goss

ypinus

Peromysc

us leuc

opus

Peromysc

us slevi

ni

Peromysc

us neso

dytes

Peromysc

us keen

i

Peromysc

us manic

ulatus

Peromysc

us polio

notus

Peromysc

us melan

otis

Peromysc

us sejug

is

Peromysc

us crinit

us

Peromysc

us califo

rnicus

Peromysc

us pember

toni

Peromysc

us canic

eps

Peromysc

us merri

ami

Peromysc

us erem

icus

Peromysc

us pseu

docrinit

us

Peromysc

us inter

parieta

lis

Peromysc

us frate

rculus

Peromysc

us guar

dia

Peromysc

us dicke

yi

Peromysc

us eva

Podomys f

loridanu

s

Neotomodo

n alston

i

Peromysc

us hoop

eri

Peromysc

us poliu

s

Peromysc

us maye

nsis

Peromysc

us mega

lops

Peromysc

us stirto

ni

Peromysc

us yuca

tanicus

Peromysc

us perfu

lvus

Peromysc

us melan

ophrys

Peromysc

us mekis

turus

Peromysc

us melan

ocarpu

s

Peromyscus m

elanurus

Peromyscus z

arhynch

us

Peromyscus m

exicanu

s

Peromyscus g

ymnotis

Peromyscus a

ztecus

Peromyscus guatem

alensis

Peromyscus grandis

Peromyscus pectoralis

Peromyscus ochraventer

Peromyscus fu

rvus

Peromyscus tru

ei

Peromyscus gratus

Peromyscus attwateri

Peromyscus nasutus

Peromyscus diffic

ilis

Peromyscus bullatus

Habromys simulatus

Habromys delica

tulus

Habromys schmidlyi

Habromys lophurus

Habromys lepturus

Habromys ixtla

ni

Habromys chinanteco

Peromyscus sa

gax

Peromyscus sp

icilegus

Peromyscus hylocetes

Peromyscus w

inkelmanni

Peromyscus beatae

Peromyscus sc

hmidlyi

Peromyscus le

vipes

Peromyscus m

adrensis

Peromyscus st

ephani

Peromyscus si

mulus

Peromyscus boylii

Phodopus roborovsk

ii

Phodopus sungorus

Phodopus campbelli

Mesocricetus auratus

Mesocricetus newtoni

Mesocricetus brandti

Mesocricetus ra

ddei

Mesocricetus ra

thgeberi

Tscherski

a triton

Cricetus cr

icetus

Cansumys canus

Allocricetulus eversm

anni

Allocricetulus cu

rtatus

Cricetulus lo

ngicaudatus

Cricetulus so

kolovi

Cricetulus m

igratorius

Cricetulus barabensis

Cricetulus altico

la

Cricetulus ka

mensis

Cricetulus la

ma

Cricetulus tib

etanus

Prometheomys schaposch

nikowi

Neofiber alleni

Ondatra zibethicus

Synaptomys borealis

Synaptomys cooperi

Myopus schistic

olor

Lemmus trimucronatus

Lemmus portenkoi

Lemmus lemmus

Lemmus sibiricu

s

Lemmus amurensis

Phenacomys intermedius

Phenacomys ungava

Arborimus pomo

Arborimus longicaudus

Arborimus albipes

Dicrostonyx n

elsoni

Dicrostonyx u

nalascensis

Dicrostonyx v

inogradovi

Dicrostonyx g

roenlandicus

Dicrostonyx n

unatakensis

Dicrostonyx t

orquatus

Dicrostonyx r

ichardsoni

Dicrostonyx h

udsonius

Hyperacrius fe

rtilis

Hyperacrius wynnei

Caryomys eva

Caryomys inez

Eothenomys melanogaster

Eothenomys cachinus

Eothenomys miletus

Eothenomys custos

Eothenomys chinensis

Eothenomys wardi

Eothenomys olitor

Eothenomys proditor

Myodes smithii

Myodes andersoni

Myodes rex

Myodes rufocanus

Myodes regulus

Myodes shanseius

Myodes glareolus

Myodes centralis

Myodes rutilus

Myodes californicus

Myodes gapperi

Alticola lemminus

Alticola stre

lzowi

Alticola argentatus

Alticola semicanus

Alticola albicaudus

Alticola roylei

Alticola baraksh

in

Alticola macrotis

Alticola sto

liczkanus

Alticola montosa

Alticola tuvinicus

Alticola olchonensis

Dinaromys bogdanovi

Pliomys lenki

Arvicola sapidus

Arvicola amphibius

Arvicola sch

erman

Ellobius lutescens

Ellobius alaicus

Ellobius talpinus

Ellobius fuscocapillus

Ellobius tancrei

Lagurus lagurus

Eolagurus luteus

Eolagurus przewalskii

Chionomys nivalis

Chionomys roberti

Chionomys gud

Lemmiscus cu

rtatus

Proedromys bedfordi

Proedromys liangshanensis

Microtus gregalis

Lasiopodomys fuscus

Lasiopodomys mandarinus

Lasiopodomys brandtii

Phaiomys leucurus

Volemys millice

ns

Volemys musseri

Neodon juldaschi

Neodon sikimensis

Neodon irene

Neodon forresti

Microtus cla

rkei

Microtus oeconomus

Microtus m

ontebelli

Microtus kik

uchii

Microtus sa

chalinensis

Microtus fo

rtis

Microtus lim

nophilus

Microtus m

iddendorffii

Microtus m

ongolicus

Microtus m

aximowiczii

Microtus m

ujanensis

Microtus evoronensis

Microtus nasarovi

Microtus agrestis

Blanfordimys bucharensis

Blanfordimys afghanus

Microtus ric

hardsoni

Microtus ca

brerae

Microtus xa

nthognathus

Microtus oregoni

Microtus longicaudus

Microtus canicaudus

Microtus townsendii

Microtus montanus

Microtus pennsylvanicus

Microtus breweri

Microtus ochrogaster

Microtus californicus

Microtus mexicanus

Microtus chrotorrhinus

Microtus umbrosus

Microtus miurus

Microtus abbreviatus

Microtus guatemalensis

Microtus oaxacensis

Microtus quasiater

Microtus pinetorum

Microtus levis

Microtus transcaspicus

Microtus arvalis

Microtus ilaeus

Microtus schelkovnikovi

Microtus dogramacii

Microtus guentheri

Microtus socialis

Microtus schidlovski

i

Microtus anatolicus

Microtus irani

Microtus paradoxus

Microtus majori

Microtus subterraneus

Microtus daghestanicus

Microtus thomasi

Microtus felteni

Microtus henseli

Microtus tatricus

Microtus hyperboreus

Microtus savii

Microtus brachycercus

Microtus qazvinensis

Microtus multiplex

Microtus liechtensteini

Microtus bavaricus

Microtus gerbei

Microtus duodecimcostatus

Microtus lusitanicus

Nyctomys s

umichrasti

Otonyctomys h

atti

Ototylomys phyllotis

Tylomys tumbalensis

Tylomys panamensis

Tylomys watsoni

Tylomys nudicaudus

Tylomys bullaris

Tylomys mirae

Tylomys fulvive

nter

Chibchanomys trichotis

Chibchanomys orcesi

Rheomys thomasi

Rheomys raptor

Rheomys mexica

nus

Rheomys underwoodi

Sigmodon alstoni

Sigmodon peruanus

Sigmodon leucotis

Sigmodon fulviventer

Sigmodon ochrognathus

Sigmodon inopinatus

Sigmodon arizonae

Sigmodon mascotensis

Sigmodon hispidus

Sigmodon toltecus

Sigmodon hirsutus

Sigmodon alleni

Sigmodon planifrons

Anotomys leander

Ichthyomys pittieri

Ichthyomys stolzmanni

Ichthyomys tweedii

Ichthyomys hydrobates

Neusticomys m

onticolus

Neusticomys o

yapocki

Neusticomys p

eruviensis

Neusticomys v

enezuelae

Neusticomys fe

rreirai

Neusticomys m

ussoi

Phaenomys ferrugineus

Rhipidomys leucodactylu

s

Rhipidomys latimanus

Rhipidomys wetzeli

Rhipidomys caucensis

Rhipidomys venezuelae

Rhipidomys austrinus

Rhipidomys cariri

Rhipidomys macconnelli

Rhipidomys ochrogaster

Rhipidomys nitela

Rhipidomys fulviventer

Rhipidomys modicus

Rhipidomys couesi

Rhipidomys venustus

Rhipidomys gardneri

Rhipidomys mastacalis

Rhipidomys emiliae

Rhipidomys macrurus

Aepeomys lugens

Aepeomys reigi

Chilomys instans

Abrawayaomys ruschii

Thomasomys apeco

Thomasomys laniger

Thomasomys rosalinda

Thomasomys incanus

Thomasomys notatus

Thomasomys niveipes

Thomasomys macrotis

Thomasomys hylophilus

Thomasomys vulcani

Thomasomys ucucha

Thomasomys pyrrhonotus

Thomasomys bombycinus

Thomasomys cinereiventer

Thomasomys erro

Thomasomys baeops

Thomasomys onkiro

Thomasomys popayanus

Thomasomys aureus

Thomasomys praetor

Thomasomys monochromos

Thomasomys paramorum

Thomasomys eleusis

Thomasomys silvestris

Thomasomys oreas

Thomasomys taczanowskii

Thomasomys caudivarius

Thomasomys cinereus

Thomasomys ischyrus

Thomasomys ladewi

Thomasomys kalinowskii

Thomasomys vestitus

Thomasomys daphne

Thomasomys rhoadsi

Thomasomys gracilis

Thomasomys cinnameus

Thomasomys hudsoni

Kunsia tomentosus

Kunsia fronto

Scapteromys aquaticus

Scapteromys tumidus

Bibimys labiosus

Bibimys chacoensis

Bibimys torresi

Lenoxus apicalis

Blarinomys breviceps

Brucepattersonius igniventris

Brucepattersonius misionensis

Brucepattersonius guarani

Brucepattersonius paradisus

Brucepattersonius soricinus

Brucepattersonius iheringi

Brucepattersonius albinasus

Brucepattersonius griserufescens

Oxymycterus delator

Oxymycterus akodontius

Oxymycterus inca

Juscelinomys guaporensis

Juscelinomys huanchacae

Juscelinomys candango

Oxymycterus amazonicus

Oxymycterus dasytrichus

Oxymycterus nasutus

Oxymycterus hiska

Oxymycterus paramensis

Oxymycterus roberti

Oxymycterus quaestor

Oxymycterus hispidus

Oxymycterus rufus

Oxymycterus hucucha

Oxymycterus angularis

Oxymycterus josei

Oxymycterus caparoae

Thalpomys cerradensis

Thalpomys lasiotis

Podoxymys roraimae

Necromys obscurus

Necromys benefactus

Necromys lactens

Necromys punctulatus

Necromys amoenus

Necromys urichi

Necromys lenguarum

Necromys lasiurus

Necromys temchuki

Thaptomys nigrita

Deltamys kempi

Akodon serrensis

Akodon bogotensis

Akodon mimus

Akodon latebricola

Akodon iniscatus

Akodon dayi

Akodon oenos

Akodon toba

Akodon molinae

Akodon dolores

Akodon siberiae

Akodon budini

Akodon simulator

Akodon varius

Akodon neocenus

Akodon albiventer

Akodon orophilus

Akodon torques

Akodon mollis

Akodon aerosus

Akodon affinis

Akodon lindberghi

Akodon surdus

Akodon cursor

Akodon montensis

Akodon reigi

Akodon paranaensis

Akodon mystax

Akodon azarae

Akodon philipmyersi

Akodon fumeus

Akodon kofordi

Akodon juninensis

Akodon aliquantulus

Akodon subfuscus

Akodon lutescens

Akodon sylvanus

Akodon pervalens

Akodon boliviensis

Akodon sanctipaulensis

Akodon spegazzinii

Akodon leucolimnaeus

Irenomys tarsalis

Euneomys chinchilloides

Euneomys mordax

Euneomys petersoni

Euneomys fossor

Reithrodon auritus

Reithrodon typicus

Rhagomys longilingua

Rhagomys rufescens

Wilfredomys oenax

Juliomys pictipes

Juliomys rimofrons

Wiedomys pyrrhorhinos

Wiedomys cerradensis

Andinomys edax

Delomys sublineatus

Delomys dorsalis

Delomys collinus

Pearsonomys annectens

Geoxus valdivianus

Chelemys macronyx

Notiomys edwardsii

Chelemys megalonyx

Chelemys delfini

Abrothrix jelskii

Abrothrix lanosus

Abrothrix longipilis

Abrothrix sanborni

Abrothrix markhami

Abrothrix olivaceus

Abrothrix andinus

Abrothrix hershkovitzi

Abrothrix illuteus

Chinchillula sahamae

Calomys sorellus

Calomys lepidus

Calomys musculinus

Calomys hummelincki

Calomys tener

Calomys laucha

Calomys expulsus

Calomys callidus

Calomys callosus

Calomys tocantinsi

Calomys venustus

Calomys fecundus

Calomys boliviae

Eligmodontia typus

Eligmodontia moreni

Eligmodontia puerulus

Eligmodontia morgani

Graomys domorum

Graomys edithae

Graomys centralis

Graomys griseoflavus

Salinomys delicatus

Andalgalomys pearsoni

Andalgalomys olrogi

Phyllotis wolffsohni

Neotomys ebriosus

Tapecomys primus

Galenomys garleppi

Punomys kofordi

Punomys lemminus

Loxodontomys micropus

Loxodontomys pikumche

Auliscomys pictus

Auliscomys boliviensis

Auliscomys sublimis

Phyllotis osilae

Phyllotis anitae

Phyllotis definitus

Phyllotis andium

Paralomys gerbillus

Phyllotis amicus

Phyllotis osgoodi

Phyllotis xanthopygus

Phyllotis bonariensis

Phyllotis limatus

Phyllotis magister

Phyllotis haggardi

Phyllotis darwini

Phyllotis caprinus

Zygodontomys brevicauda

Zygodontomys brunneus

Scolomys ucayalensis

Scolomys melanops

Transandinomys bolivaris

Transandinomys talamancae

Euryoryzomys macconnelli

Euryoryzomys emmonsae

Euryoryzomys russatus

Euryoryzomys nitidus

Euryoryzomys lamia

Euryoryzomys legatus

Noronhomys vespuccii

Oecomys rutilus

Oecomys auyantepui

Oecomys paricola

Oecomys rex

Oecomys catherinae

Oecomys cleberi

Oecomys flavicans

Oecomys superans

Oecomys bicolor

Oecomys phaeotis

Oecomys trinitatis

Oecomys mamorae

Oecomys concolor

Oecomys roberti

Oecomys speciosus

Handleyomys intectus

Handleyomys rostratus

Nephelomys albigularis

Nephelomys meridensis

Nephelomys caracolus

Nephelomys auriventer

Nephelomys devius

Nephelomys keaysi

Nephelomys levipes

Handleyomys chapmani

Handleyomys melanotis

Handleyomys alfaroi

Handleyomys saturatior

Handleyomys fuscatus

Handleyomys rhabdops

Hylaeamys yunganus

Hylaeamys oniscus

Hylaeamys megacephalus

Hylaeamys tatei

Hylaeamys laticeps

Hylaeamys perenensis

Hylaeamys acritus

Eremoryzomys polius

Mindomys hammondi

Oreoryzomys balneator

Neacomys minutus

Neacomys tenuipes

Pennatomys nivalis

Microryzomys minutus

Microryzomys altissimus

Megalomys georginae

Megalomys audreyae

Megalomys luciae

Megalomys desmarestii

Neacomys paracou

Neacomys spinosus

Neacomys dubosti

Neacomys pictus

Neacomys guianae

Neacomys musseri

Microakodontomys transitorius

Oligoryzomys microtis

Oligoryzomys eliurus

Oligoryzomys destructor

Oligoryzomys rupestris

Oligoryzomys moojeni

Oligoryzomys vegetus

Oligoryzomys fulvescens

Oligoryzomys chacoensis

Oligoryzomys nigripes

Oligoryzomys stramineus

Oligoryzomys andinus

Oligoryzomys griseolus

Oligoryzomys fornesi

Oligoryzomys longicaudatus

Oligoryzomys magellanicus

Oligoryzomys flavescens

Oligoryzomys victus

Oligoryzomys brendae

Oligoryzomys delticola

Oligoryzomys arenalis

Sooretamys angouya

Cerradomys scotti

Cerradomys subflavus

Cerradomys marinhus

Cerradomys maracajuensis

Cerradomys andersoni

Lundomys molitor

Pseudoryzomys simplex

Holochilus brasiliensis

Holochilus sciureus

Holochilus chacarius

Amphinectomys savamis

Nectomys palmipes

Nectomys rattus

Nectomys magdalenae

Nectomys apicalis

Nectomys squamipes

Oryzomys nelsoni

Oryzomys antillarum

Oryzomys palustris

Oryzomys couesi

Oryzomys curasoae

Oryzomys dimidiatus

Oryzomys gorgasi

Sigmodontomys alfari

Sigmodontomys aphrastus

Megaoryzomys curioi

Megaoryzomys Sp Now

Melanomys robustulus

Melanomys caliginosus

Melanomys zunigae

Aegialomys xanthaeolus

Aegialomys galapagoensis

Ekbletomys hypenemus

Nesoryzomys Sp B

Nesoryzomys fernandinae

Nesoryzomys narboroughi

Nesoryzomys Sp D

Nesoryzomys Sp C

Nesoryzomys swarthi

Nesoryzomys indefessus

Nesoryzomys darwini

Lophiomys imhausi

Uranomys ruddi

Deomys ferrugineus

Lophuromys woosnami

Lophuromys melanonyx

Lophuromys sikapusi

Lophuromys eisentrauti

Lophuromys medicaudatus

Lophuromys luteogaster

Lophuromys huttereri

Lophuromys rahmi

Lophuromys dieterleni

Lophuromys roseveari

Lophuromys cinereus

Lophuromys nudicaudus

Lophuromys brevicaudus

Lophuromys flavopunctatus

Lophuromys chrysopus

Acomys louisae

Acomys spinosissimus

Acomys subspinosus

Acomys wilsoni

Acomys percivali

Acomys kempi

Acomys russatus

Acomys ignitus

Acomys johannis

Acomys cineraceus

Acomys cahirinus

Acomys dimidiatus

Acomys airensis

Acomys seurati

Acomys nesiotes

Acomys minous

Acomys cilicicus

Acomys mullah

Ammodillus imbellis

Tatera indica

Gerbillurus vallinus

Gerbillurus setzeri

Desmodillus auricularis

Gerbilliscus nigricaudus

Gerbilliscus robustus

Gerbillurus paeba

Gerbillurus tytonis

Gerbilliscus guineae

Gerbilliscus kempi

Gerbilliscus inclusus

Gerbilliscus gambiana

Gerbilliscus boehmi

Gerbilliscus brantsii

Gerbilliscus afra

Gerbilliscus leucogaster

Gerbilliscus validus

Gerbilliscus phillipsi

Pachyuromys duprasi

Desmodilliscus braueri

Taterillus lacustris

Taterillus gracilis

Taterillus emini

Taterillus arenarius

Taterillus congicus

Taterillus pygargus

Taterillus petteri

Taterillus tranieri

Psammomys obesus

Psammomys vexillaris

Brachiones przewalskii

Rhombomys opimus

Meriones hurrianae

Meriones tamariscinus

Meriones persicus

Meriones rex

Meriones zarudnyi

Meriones sacramenti

Meriones meridianus

Meriones chengi

Meriones dahli

Meriones crassus

Meriones shawi

Meriones grandis

Meriones libycus

Meriones vinogradovi

Meriones tristrami

Meriones arimalius

Meriones unguiculatus

Sekeetamys calurus

Microdillus peeli

Gerbillus campestris

Gerbillus rupicola

Gerbillus somalicus

Gerbillus dasyurus

Gerbillus jamesi

Gerbillus maghrebi

Gerbillus lowei

Gerbillus stigmonyx

Gerbillus bottai

Gerbillus mackillingini

Gerbillus harwoodi

Gerbillus simoni

Gerbillus percivali

Gerbillus pusillus

Gerbillus mesopotamiae

Gerbillus watersi

Gerbillus juliani

Gerbillus grobbeni

Gerbillus muriculus

Gerbillus nanus

Gerbillus principulus

Gerbillus brockmani

Gerbillus henleyi

Gerbillus poecilops

Gerbillus amoenus

Gerbillus famulus

Gerbillus andersoni

Gerbillus gleadowi

Gerbillus cosensis

Gerbillus cheesmani

Gerbillus hesperinus

Gerbillus pyramidum

Gerbillus burtoni

Gerbillus agag

Gerbillus acticola

Gerbillus occiduus

Gerbillus pulvinatus

Gerbillus tarabuli

Gerbillus perpallidus

Gerbillus nancillus

Gerbillus hoogstraali

Gerbillus rosalinda

Gerbillus dunni

Gerbillus gerbillus

Gerbillus nigeriae

Gerbillus latastei

Gerbillus aquilus

Gerbillus floweri

Phloeomys cumingi

Phloeomys pallidus

Carpomys phaeurus

Carpomys melanurus

Batomys salomonseni

Batomys dentatus

Batomys granti

Batomys russatus

Crateromys paulus

Crateromys australis

Crateromys heaneyi

Crateromys schadenbergi

Micromys minutus

Maxomys inas

Maxomys pagensis

Maxomys whiteheadi

Maxomys hylomyoides

Maxomys inflatus

Sommeromys macrorhinos

Crunomys fallax

Crunomys celebensis

Crunomys suncoides

Crunomys melanius

Maxomys bartelsii

Maxomys panglima

Maxomys wattsi

Maxomys hellwaldii

Maxomys ochraceiventer

Maxomys baeodon

Maxomys dollmani

Maxomys rajah

Maxomys alticola

Maxomys moi

Maxomys surifer

Maxomys musschenbroekii

Hadromys humei

Hadromys yunnanensis

Dacnomys millardi

Anonymomys mindorensis

Leopoldamys neilli

Leopoldamys sabanus

Leopoldamys siporanus

Leopoldamys milleti

Leopoldamys edwardsi

Leopoldamys ciliatus

Chiromyscus chiropus

Tonkinomys daovantieni

Niviventer langbianis

Niviventer hinpoon

Niviventer cremoriventer

Niviventer rapit

Niviventer cameroni

Niviventer fraternus

Niviventer andersoni

Niviventer excelsior

Niviventer lepturus

Niviventer culturatus

Niviventer confucianus

Niviventer tenaster

Niviventer coninga

Niviventer brahma

Niviventer eha

Niviventer fulvescens

Niviventer niviventer

Lenothrix canus

Eropeplus canus

Lenomys meyeri

Pithecheirops otion

Pithecheir melanurus

Pithecheir parvus

Melasmothrix naso

Tateomys macrocercus

Tateomys rhinogradoides

Echiothrix centrosa

Echiothrix leucura

Margaretamys parvus

Margaretamys beccarii

Margaretamys elegans

Srilankamys ohiensis

Saxatilomys paulinae

Berylmys mackenziei

Berylmys manipulus

Berylmys berdmorei

Berylmys bowersi

Bullimus luzonicus

Bullimus gamay

Bullimus bagobus

Sundamys infraluteus

Sundamys muelleri

Sundamys maxi

Palawanomys furvus

Paruromys dominator

Papagomys theodorverhoeveni

Papagomys armandvillei

Komodomys rintjanus

Bunomys chrysocomus

Bunomys fratrorum

Bunomys prolatus

Bunomys coelestis

Bunomys penitus

Bunomys andrewsi

Nesokia indica

Nesokia bunnii

Kadarsanomys sodyi

Bandicota bengalensis

Bandicota indica

Bandicota savilei

Diplothrix legata

Taeromys callitrichus

Taeromys microbullatus

Taeromys punicans

Taeromys taerae

Taeromys hamatus

Taeromys celebensis

Taeromys arcuatus

Rattus fuscipes

Rattus lutreolus

Rattus timorensis

Rattus tunneyi

Rattus hainaldi

Rattus sordidus

Rattus villosissimus

Rattus colletti

Rattus omichlodes

Rattus richardsoni

Rattus niobe

Rattus arfakienis

Rattus pococki

Rattus arrogans

Rattus hoogerwerfi

Rattus nativitatis

Rattus leucopus

Rattus mordax

Rattus sanila

Rattus feliceus

Rattus giluwensis

Rattus jobiensis

Rattus steini

Rattus elaphinus

Rattus novaeguineae

Rattus vandeuseni

Rattus verecundus

Rattus stoicus

Rattus praetor

Rattus pelurus

Rattus salocco

Rattus xanthurus

Rattus bontanus

Rattus marmosurus

Rattus everetti

Rattus pyctoris

Abditomys latidens

Tarsomys apoensis

Tarsomys echinatus

Limnomys bryophilus

Limnomys sibuanus

Tryphomys adustus

Paulamys naso

Nesoromys ceramicus

Rattus satarae

Rattus norvegicus

Rattus nitidus

Rattus annandalei

Rattus exulans

Rattus andamanensis

Rattus tawitawiensis

Rattus argentiventer

Rattus mindorensis

Rattus hoffmanni

Rattus montanus

Rattus losea

Rattus macleari

Rattus rattus

Rattus tanezumi

Rattus mollicomulus

Rattus korinchi

Rattus ranjiniae

Rattus baluensis

Rattus morotaiensis

Rattus osgoodi

Rattus koopmani

Rattus enganus

Rattus blangorum

Rattus adustus

Rattus lugens

Rattus tiomanicus

Rattus palmarum

Rattus burrus

Rattus simalurensis

Haeromys pusillus

Haeromys margarettae

Haeromys minahassae

Hapalomys delacouri

Hapalomys longicaudatus

Chiropodomys muroides

Chiropodomys major

Chiropodomys gliroides

Chiropodomys calamianensis

Chiropodomys pusillus

Chiropodomys karlkoopmani

Apomys datae

Apomys abrae

Apomys littoralis

Apomys musculus

Apomys microdon

Apomys sacobianus

Apomys camiguinensis

Apomys hylocetes

Apomys insignis

Apomys gracilirostris

Archboldomys musseri

Archboldomys luzonensis

Archboldomys kalinga

Rhynchomys isarogensis

Rhynchomys tapulao

Rhynchomys banahao

Rhynchomys soricoides

Chrotomys silaceus

Chrotomys sibuyanensis

Chrotomys gonzalesi

Chrotomys whiteheadi

Chrotomys mindorensis

Lorentzimys nouhuysi

Anisomys imitator

Xenuromys barbatus

Macruromys major

Macruromys elegans

Chiruromys forbesi

Chiruromys vates

Chiruromys lamia

Hyomys goliath

Hyomys dammermani

Coryphomys buehleri

Coryphomys musseri

Spelaeomys florensis

Pogonomys macrourus

Pogonomys sylvestris

Pogonomys championi

Pogonomys loriae

Pogonomys fergussoniensis

Mammelomys lanosus

Mammelomys rattoides

Mallomys gunung

Mallomys aroaensis

Mallomys rothschildi

Mallomys istapantap

Abeomelomys sevia

Pogonomelomys mayeri

Pogonomelomys bruijnii

Pogonomelomys brassi

Parahydromys asper

Baiyankamys habbema

Baiyankamys shawmayeri

Hydromys hussoni

Hydromys ziegleri

Hydromys neobritannicus

Hydromys chrysogaster

Crossomys moncktoni

Coccymys albidens

Coccymys ruemmleri

Paraleptomys rufilatus

Paraleptomys wilhelmina

Leptomys signatus

Leptomys elegans

Leptomys ernstmayri

Xeromys myoides

Microhydromys musseri

Microhydromys richardsoni

Pseudohydromys murinus

Pseudohydromys occidentalis

Pseudohydromys ellermani

Pseudohydromys fuscus

Pseudohydromys germani

Zyzomys woodwardi

Zyzomys pedunculatus

Zyzomys argurus

Zyzomys maini

Zyzomys palatalis

Leggadina forresti

Leggadina lakedownensis

Notomys fuscus

Notomys macrotis

Notomys mordax

Notomys alexis

Notomys aquilo

Notomys mitchellii

Notomys longicaudatus

Notomys cervinus

Notomys amplus

Mastacomys fuscus

Pseudomys higginsi

Pseudomys fieldi

Pseudomys glaucus

Pseudomys australis

Pseudomys hermannsburgensis

Pseudomys occidentalis

Pseudomys delicatulus

Pseudomys desertor

Pseudomys novaehollandiae

Pseudomys pilligaensis

Pseudomys oralis

Pseudomys shortridgei

Pseudomys bolami

Pseudomys gouldii

Pseudomys nanus

Pseudomys gracilicaudatus

Pseudomys fumeus

Pseudomys apodemoides

Pseudomys albocinereus

Pseudomys calabyi

Pseudomys patrius

Pseudomys johnsoni

Pseudomys chapmani

Leporillus conditor

Leporillus apicalis

Conilurus penicillatus

Conilurus albipes

Mesembriomys gouldii

Mesembriomys macrurus

Uromys porculus

Uromys imperator

Uromys caudimaculatus

Uromys siebersi

Uromys rex

Uromys anak

Uromys hadrourus

Uromys neobritannicus

Uromys emmae

Uromys boeadii

Protochromys fellowsi

Paramelomys moncktoni

Paramelomys rubex

Paramelomys steini

Paramelomys platyops

Paramelomys lorentzii

Paramelomys naso

Paramelomys levipes

Paramelomys gressitti

Paramelomys mollis

Solomys ponceleti

Solomys sapientis

Solomys salamonis

Solomys spriggsarum

Solomys salebrosus

Melomys bougainville

Melomys aerosus

Melomys lutillus

Melomys howi

Melomys cervinipes

Melomys matambuai

Melomys talaudium

Melomys fulgens

Melomys arcium

Melomys caurinus

Melomys leucogaster

Melomys rubicola

Melomys obiensis

Melomys fraterculus

Melomys burtoni

Melomys frigicola

Melomys bannisteri

Melomys capensis

Melomys dollmani

Melomys rufescens

Melomys cooperae

Melomys paveli

Diomys crumpi

Cremnomys cutchicus

Cremnomys elvira

Madromys blanfordi

Millardia meltada

Millardia kondana

Millardia kathleenae

Millardia gleadowi

Otomys denti

Otomys cuanzensis

Otomys lacustris

Otomys anchietae

Parotomys littledalei

Parotomys brantsii

Otomys barbouri

Otomys unisulcatus

Otomys sloggetti

Otomys irroratus

Otomys saundersiae

Otomys laminatus

Otomys angoniensis

Otomys burtoni

Otomys occidentalis

Otomys tropicalis

Otomys typus

Golunda ellioti

Lamottemys okuensis

Oenomys hypoxanthus

Oenomys ornatus

Thamnomys venustus

Thamnomys kempi

Thamnomys schoutedeni

Stochomys longicaudatus

Dephomys defua

Hybomys basilii

Hybomys lunaris

Hybomys univittatus

Hybomys planifrons

Hybomys trivirgatus

Hybomys badius

Aethomys kaiseri

Aethomys hindei

Aethomys bocagei

Aethomys ineptus

Aethomys chrysophilus

Aethomys stannarius

Aethomys thomasi

Aethomys nyikae

Aethomys silindensis

Aethomys namaquensis

Aethomys granti

Grammomys gigas

Grammomys ibeanus

Grammomys dolichurus

Grammomys caniceps

Grammomys cometes

Grammomys aridulus

Grammomys kuru

Grammomys buntingi

Grammomys macmillani

Grammomys minnae

Grammomys dryas

Malpaisomys insularis

Canariomys bravoi

Canariomys tamarani

Thallomys loringi

Thallomys paedulcus

Thallomys nigricauda

Thallomys shortridgei

Dasymys incomtus

Dasymys rufulus

Dasymys montanus

Dasymys nudipes

Dasymys foxi

Desmomys harringtoni

Desmomys yaldeni

Mylomys dybowskii

Mylomys rex

Pelomys fallax

Pelomys hopkinsi

Pelomys minor

Pelomys campanae

Pelomys isseli

Rhabdomys pumilio

Arvicanthis nairobae

Arvicanthis neumanni

Arvicanthis blicki

Arvicanthis abyssinicus

Arvicanthis rufinus

Arvicanthis niloticus

Arvicanthis ansorgei

Lemniscomys mittendorfi

Lemniscomys striatus

Lemniscomys barbarus

Lemniscomys zebra

Lemniscomys rosalia

Lemniscomys linulus

Lemniscomys roseveari

Lemniscomys hoogstraali

Lemniscomys griselda

Lemniscomys macculus

Lemniscomys bellieri

Leimacomys buettneri

Vernaya fulva

Vandeleuria nolthenii

Vandeleuria nilagirica

Vandeleuria oleracea

Myomyscus yemeni

Myomyscus brockmani

Myomyscus angolensis

Stenocephalemys albocaudata

Stenocephalemys griseicauda

Stenocephalemys albipes

Stenocephalemys ruppi

Praomys delectorum

Praomys morio

Mastomys pernanus

Mastomys shortridgei

Heimyscus fumosus

Hylomyscus denniae

Hylomyscus stella

Hylomyscus parvus

Hylomyscus alleni

Hylomyscus baeri

Hylomyscus carillus

Hylomyscus aeta

Hylomyscus grandis

Myomyscus verreauxii

Zelotomys hildegardeae

Zelotomys woosnami

Colomys goslingi

Nilopegamys plumbeus

Mastomys kollmannspergeri

Mastomys huberti

Mastomys coucha

Mastomys natalensis

Mastomys erythroleucus

Mastomys awashensis

Praomys verschureni

Praomys lukolelae

Praomys degraaffi

Praomys minor

Praomys jacksoni

Praomys derooi

Praomys daltoni

Praomys petteri

Praomys hartwigi

Praomys obscurus

Praomys rostratus

Praomys tullbergi

Praomys misonnei

Praomys mutoni

Tokudaia muenninki

Tokudaia tokunoshimensis

Tokudaia osimensis

Apodemus argenteus

Apodemus speciosus

Apodemus agrarius

Apodemus chevrieri

Apodemus peninsulae

Apodemus latronum

Apodemus semotus

Apodemus draco

Apodemus gurkha

Apodemus mystacinus

Apodemus epimelas

Apodemus pallipes

Apodemus uralensis

Apodemus witherbyi

Apodemus alpicola

Apodemus sylvaticus

Apodemus hyrcanicus

Apodemus flavicollis

Apodemus ponticus

Apodemus rusiges

Malacomys edwardsi

Malacomys longipes

Malacomys cansdalei

Muriculus imberbis

Mus pahari

Mus vulcani

Mus crociduroides

Mus mayori

Mus saxicola

Mus platythrix

Mus fernandoni

Mus phillipsi

Mus shortridgei

Mus caroli

Mus cookii

Mus cervicolor

Mus famulus

Mus terricolor

Mus booduga

Mus fragilicauda

Mus spretus

Mus musculus

Mus spicilegus

Mus macedonicus

Mus cypriacus

Mus sorella

Mus baoulei

Mus bufo

Mus triton

Mus setulosus

Mus mahomet

Mus orangiae

Mus goundae

Mus mattheyi

Mus haussa

Mus oubanguii

Mus indutus

Mus callewaerti

Mus musculoides

Mus setzeri

Mus minutoides

Mus tenellus

Mus neavei

a

d

e

cb

h

g

f

i

Time (Millions of years ago)

Petrogale persephone

Petrogale burbidgei

Petrogale concinna

Petrogale brachyotis

Petrogale xanthopus

Petrogale coenensis

Petrogale lateralis

Petrogale godmani

Petrogale rothschildi

Petrogale purpureicollis

Petrogale herberti

Petrogale penicillata

Petrogale assimilis

Petrogale sharmani

Petrogale inornata

Petrogale mareeba

a)

b)

c)

Figure 3 Bioregion map and phylogenetic tree of world mammals with ancestral range reconstruction, generated with Infomap Bioregions.a) Phylogenetic tree of 5,747 mammals computed from (Faurby and Svenning, 2015), fully zoomable on the online application. Ancestralranges reconstructed under Fitch parsimony. Pie charts depict most parsimonious ancestral ranges at nodes, and current distributions forextant species. Branch lines are scaled to the number of terminals subtending each branch, in order to improve visualisation of the overalltree structure. b) Magnified part of the tree, highlighting the rock-wallabies (genus Petrogale) which are currently distributed across severalbioregions in Australia. This analysis suggests that all rock-wallabies, including the Yellow-footed rock-wallaby (Petrogale xanthopus), whichis the most indicative species of the southeast bioregion (i), originated from a common ancestor in northern Australia. c) Bioregion map ofworld mammals using species point occurrences from GBIF. White areas have insufficient data and were excluded from the analysis. Coloursare used consistently across the subfigures. Table 2 shows information about labelled bioregions.

2001), see Figure 4. For these analyses we used the mapcurvesalgorithm (Hargrove et al., 2006) as implemented by (vanLoon, 2006). Mapcurves is a quantitative method to comparethe spatial concordance between categorical maps, by calcu-lating a goodness of fit (GOF) for each polygon in a map ofinterest based on the degree of spatial overlap with the poly-gons of a reference map. The results can be summarized in a

global GOF score. Mapcurves is resolution independent, doesnot require the same number of categories in both maps, andany polygon in a map that can be exactly comprised of a setof polygons in the reference map will show a perfect fit. How-ever, the algorithm generally indicates a poor fit when a finerresolution map is compared to a coarser map. A limitation ofthis asymmetry becomes apparent when the map of interest has

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Table 2 Highlighted mammalian bioregions, sorted on species richness. For exact locations, the indices (a)–(i) are displayed on the bioregionmap in Figure 3. Common names from Encyclopedia of Life at http://eol.org

Location Records Species Cells Most common species (records) Most indicative species (score)(a) South America 69,757 1,448 254 Glossophaga soricina (1,588) Uroderma bilobatum (36)

Pallas’s long-tongued bat Tent-making Bat(b) Africa 38,258 1,105 323 Mastomys natalensis (991) Mus musculoides (58)

Common African rat Kasai mouse(c) Malay Archipelago 8,788 576 103 Rattus exulans (370) Ptenochirus jagori (155)

Polynesian rat Greater Musky Fruit Bat(d) North America 279,416 812 426 Peromyscus maniculatus (17,600) Thomomys bottae (3.3)

Deer mouse Valley Pocket Gopher(e) Andes 4,709 394 46 Phyllotis xanthopygus (280) Akodon albiventer (189)

Yellow-rumped leaf-eared mouse White-bellied grass mouse(f) Europe 635,148 389 266 Meles meles (46,299) Talpa europaea (1.2)

Eurasian badger European mole(g) New Guinea 5,107 325 46 Syconycteris australis (260) Echymipera kalubu (220)

Southern blossom bat Common Echymipera(h) SE Australia 298,374 269 39 Phascolarctos cinereus (26,029) Petaurus australis (2.2)

Koala Yellow-bellied glider(i) SE Australia 43,669 147 22 Macropus robustus (9,176) Petrogale xanthopus (6.2)

Hill wallaroo Yellow-footed rock-wallaby

coarser resolution than the chosen reference map in some ormost of the areas, but finer resolution in other areas. Therefore,the globally best GOF map may have areas of finer resolutionwhere the local GOF is poor, whereas the bioregions in thereference map covering the same area may have better localGOF. See Hargrove et al. (2006) for a detailed description ofthe method.

The results of the comparison show a generally good fitbetween the bioregions identified by Infomap Bioregions andthe WWF ecoregions. The fit was better for amphibians (globalGOF 0.65) than for mammals (0.54), which might be relatedto the the data used for map creation (range polygons vs. pointoccurrences, respectively). The spatial visualization of theGOF scores shows that the fit is very good for most areas.Differences are mainly associated with a number of very smallbioregions, mostly in the Andes for amphibians and mostly inAsia and northern Africa for mammals. For mammals, many ofthe small bioregions recognized by Infomap Bioregions seemto derive from low data availability. A low fit is also partly anartefact of the asymmetry of the measure as mentioned above,especially for amphibians in the Andes.

We also compared the bioregions identified by InfomapBioregions to the zoogeographic regions from (Holt et al.,2013), which for the amphibians were based on approximatelythe same data, but delimited using a beta-diversity method in-cluding phylogenetic information. The GOF was overall verygood, with a global GOF score of 0.77 for amphibians and0.52 for mammals (see Supplementary Material). In summary,the results of the pairwise comparisons show that bioregionsdelimited by Infomap Bioregions generally correspond well tocommonly used bioregionalization maps, despite differencesin the underlying data and methodology applied.

C O N C L U S I O N S

Designed to make data-driven identification of bioregions sim-ple and effective, we introduced the web application InfomapBioregions and demonstrated its flexibility. A user can loadspecies data from both point occurrences and range polygons,modify parameters directly in the web interface, and exportresults to various formats for high-quality printing or furtherbiogeographical analyses. The web application uses adaptivespatial resolution, can process millions of records in a few min-utes, and applies bipartite network clustering that outperformstraditional methods based on similarity indices. Moreover, theuser can load phylogenetic data for the species and explorehow the bioregions map to the phylogenetic tree. We validatedthe application on two large datasets of amphibians and mam-mals and anticipate that Infomap Bioregions will become astandard tool in many studies in ecology, evolution, conserva-tion biology and historical biogeography.

AVA I L A B I L I T Y A N D F O R T H C O M I N G E X T E N S I O N S

Infomap Bioregions is made open source under the GNUAGPL v3+ license. It is written in JavaScript and buildson a set of open source libraries (see dependencies inpackage.json). Because it is a pure client-side application,all data stay and all calculations run on the user’s computer.Moreover, all heavy calculations run in a background thread.Together, this means improved privacy and performance.

Infomap Bioregions is available at http://bioregions.mapequation.org and the source code is freely available athttp://github.com/mapequation/bioregions.

Possible forthcoming extensions include batch runs, addi-tional methods to find indicator species and bioregions, hierar-chical clustering of bioregions, deeper integration of phyloge-netic information and significance clustering with bootstrap to

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a) Infomap amphibians b) Infomap mammals

c) WWF ecoregions d) mapcurves

Goodness of fit (GOF)

% o

f b

iore

gio

ns w

ith h

igh

er

GO

F

Amphibians

Mammals

e) Amphibians GOF 0.65 f) Mammals GOF 0.54

0.00 0.25 0.50 0.75 1.00GOF

Infomap bioregions compared to WWF ecoregions

Figure 4 Comparison between Infomap bioregions and WWF ecoregions using the Mapcurves algorithm (Hargrove et al., 2006). a) Infomapbioregions for the amphibian dataset; b) Infomap bioregions for the mammalian dataset; c) WWF ecoregions from (Olson et al., 2001); d)Mapcurves as a measure for the goodness of fit (GOF) for the Infomap bioregions with respect to the WWF ecoregions. The graph showsthe percentage of bioregions with a GOF score better than the corresponding value on the horizontal axis (zero to one). A perfect fit for allbioregions would be indicated by a horizontal line at the top; e) GOF map of the Infomap bioregions for amphibians and the total GOF score; f)GOF map of the Infomap bioregions for mammals and the total GOF score. The fit of the bioregions to the WWF ecoregions is generally verygood, with the exception of several very small bioregions identified by Infomap Bioregions.

find which bioregional boundaries are statistically significant.The authors welcome suggestions for enhancements.

S U P P L E M E N TA RY M AT E R I A L S

Data available from the Dryad Digital Repository:http://dx.doi.org/10.5061/dryad.2s201

A C K N O W L E D G M E N T S

We thank Daril Vilhena, Shawn Laffan, and our colleagues andstudents for discussions. We also thank Anna EklÃuf and twoanonymous reviewers, associate editor James Albert, and chiefeditors Frank Andersson and Thomas Near for constructivecomments on this manuscript.

F U N D I N G

This work was supported by the Swedish Research Council(B0569601 to A.A. and 2012-3729 to M.R.); the EuropeanResearch Council under the European UnionâAZs SeventhFramework Programme (FP/2007-2013 and ERC Grant Agree-ment n. 331024 to A.A.); a Wallenberg Academy Fellowshipto A.A.; and the São Paulo Research Foundation (2013/04170-8 and 2014/18837-7 to T.B.G.).

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