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金光閃閃瑞氣千條 - 視覺系統 焦傳金 國立清華大學生命科學系

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Page 1: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

金光閃閃瑞氣千條 - 視覺系統

焦傳金

國立清華大學生命科學系

Page 2: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Are they still or moving?

Page 3: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

http://web.mit.edu/persci/people/adelson/checkershadow_illusion.html

The squares marked A and B are the same shade of gray

Page 4: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

The original image plus two stripes.

By joining the squares marked A and B

with two vertical stripes of the same

shade of gray, it becomes apparent

that both squares are the same.

Page 5: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Take home messages

1. Your retina is NOT like your digital camera.

2. What your eye see is NOT exactly what you actually see.

Page 6: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Our retina is “inversed”, and we have a “blind spot”

Page 7: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

視蛋白(o p s i n)與視黃醛(retinal,非常類似維生素A)

Page 8: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

視色素一旦吸收了光線,新增加的能量就會改變視黃醛的形狀

Page 9: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

視色素一旦吸收了光線,引發一連串的分子事件,造成錐細胞的興奮

Page 10: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

To see the back of human eye

Ophthalmoscope

Page 11: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Fundus of human eye

Page 12: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Find your blind spot and beyond

Page 13: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 14: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

What if the backgrounds on both sides are different?

Page 15: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 16: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

What if we reverse the green and the yellow background?

Page 17: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 18: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

What if it is more than a spot?

Page 19: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 20: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Finally, what if it is not a plain background?

Page 21: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 22: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

What your eye see is not exactly what you

actually see

Page 23: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Contrast between an object and its background

Page 24: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Nobel Laureate 1967

Page 25: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Lateral inhibition

Page 26: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 27: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 28: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

On-center, off-surround cells

Responds to light in the center and inhibited by light in

the surround. Inhibited by a dark spot in the center and

excited by dark in the surround

Page 29: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Your retina is NOT like your digital camera

Page 30: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 31: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 33: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Orientation columns

in the visual cortex of

the monkey

The primary visual cortex

is organized into

functional modules

9x12 mm

Pinwheel-like area

Page 34: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 35: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Using optical imaging the form of the orientation columns

in the cortex for all possible orientations takes on the

shape of small pinwheels

Page 36: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Parallel pathways convey information from the retina to parietal

and temporal cortices

Page 37: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Motion and color are processed in different brain areas

Page 38: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Monocular cues create far-field depth perception

Occlusion (4,5)

Linear perspective (6/7, 8/9)

Size perspective (1,2)

Familiar size (2,3)

Optic flow: the full-field motion that results

from the observer’s own movement

Page 39: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Neural basis of stereoscopic vision

Binocular disparity

Page 40: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Cells are binocular and receive input from corresponding

visual fields.

movie

Page 41: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Random dot stereograms separate stereopsis from object vision

See this pair through a

stereoscope, or by training

the eyes to focus outside

the image plane

Ocular disparity, rather

than the form

Page 42: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Cells in V2 respond to both illusory and actual contours

Illusory contour

Page 43: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Cells in V4 respond to form, many inferior temporal neurons

also respond to color

A “Circle” selective cell A “pink” color selective cell

Page 44: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Recognition of faces and other complex forms depend upon the

inferior temporal cortex

A “face” selective cell

Page 45: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 46: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 47: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 48: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Size constancy

Page 49: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 50: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Figure-ground recognition

Page 51: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Young lady & old lady

Page 52: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 53: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Color blind

1. What do color blind people see?

2. Can your slides been seen by color blind people?

Page 54: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

In its most severe forms, color

blindness is caused by the absence of

one of the cone visual pigments.

Shown here, the spectral sensitivities

of the cone pigments in color normal

trichromats are compared with those

of a color blind person. Also compare

the spectrum as it appears to a color

normal person with the illustration of

how it might look to a red-green color

blind person.

Page 55: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

1 2

3 4

1 2

3 4

1 2

3 4

Page 56: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 57: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 58: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 59: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 60: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual
Page 61: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

2008/3/28

Page 62: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

缺少紅色感光細胞

(紅綠色盲)

2008/3/28

http://www.vischeck.com/

Page 63: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

缺少綠色感光細胞

(紅綠色盲)

2008/3/28

http://www.vischeck.com/

Page 64: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

缺少藍色感光細胞

(黃藍色盲)

2008/3/28

http://www.vischeck.com/

Page 65: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

The difficulties with certain

colors can be illustrated on a

color wheel. For a severely red-

green color blind person there

are only two hues, the ones a

color normal person sees as

yellow and blue. Intermediate

colors, the ones seen as blueish-

green and magenta, appear gray.

Page 66: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Clear graphics are very important and often the source of the most difficulty;

here are two examples showing the difference between good and bad use of

color. The Trick is to keep brightness differences large and to avoid color

combinations that do not contrast well.

Page 67: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

A comparison of the two color wheels shows which color combinations would be

difficult to see. Graphics cause the most problems, but the colors can be eco-

nomically illustrated as text examples, below. The right-hand column illustrates

how the left hand column might look to a color blind person.

Page 68: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual

Some examples of color combinations that are easily seen by a color blind person

are shown below. These are illustrated as text examples but these principles are

most important to keep in mind when preparing drawings, graphs, and figures.

Page 69: 金光閃閃瑞氣千條 視覺系統 - National Tsing Hua Universitylife.nthu.edu.tw/~bioclass/files/990424-1.pdf · 2011-11-14 · Cells in V2 respond to both illusory and actual