flores villela et al (2008)-sceloporus ochoterenae.pdf

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Catalogue of American Amphibians and Reptiles.

Flores Villela, O., H.M. Smith, E.A. Liner, and D.Chiszar. 2008. Sceloporus ochoterenae.

Sceloporus ochoterenaeSmith

Ochoterenas Spiny Lizard

Sceloporus scalaris: Dumril, Bocourt, and Moc-quard 1874:202 (nec Wiegmann) (part).

Sceloporus gratiosus: Ferrari-Perez 1886:193 (necYarrow).

Sceloporus aeneusvar.: Ferrari-Perez 1886:193 (necWiegmann).

Sceloporus graciosus: Cope 1887:36 (part).Sceloporus variabilis: Gnther 1890:75 (part).Sceloporus ochoterenSmith 1934:269. Type-local-

ity, "two miles north of Mazatln (12 miles south ofChilpancingo), Guerrero, Mexico". Holotype, EHT

/HMS 1075, Field Museum of Natural History(FMNH) 100052, male, collected 26 June 1932 by

E.H. Taylor and H.M. Smith.Sceloporus aeneus: Ahl 1934:184 (nec Wiegmann).Sceloporus ochoterenai: Smith and Taylor 1950b:

133.Lysoptychus ochoterenae: Larsen and Tanner 1975:

18.Sceloporus ochoterenaei: Wiens 1993:294. Lapsus.

CONTENT. No subspecies are currently recog-nized.

DEFINITION. Maximum SVL ca. 55 mm.; about 10paravertebral, transverse dark spots on each side ofbody, separated by a brownish vertebral line, and

bordered by a dorsolateral light line, in females andyoung males; adult males may become nearly uni-form tan above, and always have a weaker dorsalpattern than females; head scales more or less nor-mal, except for absence of postrostrals; internasalsand nasals contacting rostral; a single row of largesupraoculars; dorsal scales 3846, mean 42.4, keel-ed, mucronate, weakly denticulate; lateral scales sim-ilar in character to dorsals, half or one third as large;ventral scales smaller than laterals, smallest medial-ly, smooth, notched except for preanals; femoralpores 1016 on each side, the two series separatedby 26 scales; preanals keeled in females; a pair ofenlarged postanals in males. Oviparous.

DIAGNOSIS. This species is one of the most dis-tinctive of the genus, differing from all others by thecombination of absence of postrostrals and post-femoral dermal pockets, and presence of 1016 fe-moral pores on each side. The keeled preanals infemales are very unusual, otherwise found only in thesiniferusspecies group (to which S. ochoterenaewasonce assigned [Smith 1939] on this basis). The onlyother species with no postrostrals and no postfemoraldermal pocket is S. jalapae, its closest relative, withwhich it forms a separatejalapaespecies group (Tho-mas and Dixon 1976, Cole 1978, Wiens and Reeder

1997). S. ochoterenae differs from S. jalapae mostconspicuously in number of femoral pores (1016, vs1721) and number of dorsals, occiput to rear bor-ders of thighs (3846, vs 5062). The only other spe-

cies of the genus without postrostrals is S. maculo-sus, which has postfemoral dermal pockets and dif-fers from the jalapae species group in numerousother respects.

DESCRIPTIONS. Other descriptions are in Cas-tro-Franco and Aranda-Escobar (1984), HernndezGarca (1989), Khler and Heimes (2002), Saldaade la Riva and Prez Ramos (1987), and Smith(1934, 1939, 1984).

ILLUSTRATIONS. A color photograph of a maleand a female is in Khler and Heimes (2002), andhead scales are illustrated in a line drawing by Smith

(1934, 1939).

DISTRIBUTION. Observed in summer, autumnand winter in dry situations on the ground in rockyareas in deciduous and juniper forests at elevationsfrom 520 to reputedly 2134 m northward from south-eastern central Guerrero (Rincn/Cajones) to centralMorelos (Cuernavaca region), westward into extremesouthern Mexico State, eastward to western Puebla,and in Guerrero to near its northeastern border withOaxaca at Alpoyeca (Guerrero range mapped by Sal-daa de la Riva and Prez Ramos, 1987). Thespecies undoubtedly occurs in adjacent areas in thestate of Oaxaca.

The material reported as Sceloporus scalaris byGadow (1905) from Tierra Colorada, Guerrero, as-signed to this species by Smith (1939), has incorrectlocality data (Davis and Dixon 1961, Edmundo Prez-Ramos, pers. comm.), but its identity as Sceloporusochoterenaehas been confirmed (Smith et al., 2000).Gadow's other records in Guerrero for scalaris(Chil-pancingo, Ro Balsas) are almost certainly correctlyassigned to ochoterenae. The specimen from Coyoa-cn, Distrito Federal, reported by Smith (1939) hasincorrect locality data. Olson et al. (1986) listed thespecies for Puente de Ixtla, Veracruz; the state was alapsus for Morelos. Smith (1939) erroneously refer-

REPTILIA: SQUAMATA: PHRYNOSOMATIDAE Sceloporus ochoterenae

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FIGURE 1. A female S. ochoterenae from Ixcateopan de

Cuauhtemoc, Guerrero. Photograph by Luis Canseco

Mrquez.


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red Copes (1887) and Ferrari-Perezs (1886) recordsfor [Ixtlan de] Matamoros to S. jalapae. Records forOaxaca (Casas Andreu et al. 1997) pertain to S. jala-

pae.

FOSSIL RECORD. None. The name was men-tioned by Wellstead (1982) in connection with fossilsof other species of the genus in Nebraska.

PERTINENT LITERATURE. Anatomy (Larsenand Tanner 1974, 1975; Olson et al. 1986, 1987), be-havior (Purdue and Carpenter 1972), bibliogra-phies (Smith and Smith 1976, 1993), checklists,keys, and similar compendia (Aguilar Bentez1990; Bell et al. 2003; Casas Andreu 1982; CasasAndreu et al. 2004; Castro Franco et al. 1986; Coch-ran 1961; Cuesta Terron 1932; Duellman 1965;

Flores Villela 1993; Flores Villela et al. 1995; Frankand Ramus 1995; Hutchins et al. 2003; Liner 1994,1996, 2007; Marx 1976; Smith 1936, 1987, 1991;Smith and Taylor 1950a,b; Smith et al. 1964; Sokolof1988; Taylor 1944), conservation (Bojrquez-Tapiaet al. 1995; Garca 2006), ecology and zoogeogra-phy (Camarillo and Aguilar 1992; Camarillo andSmith 1992; Castro Franco 2002; Castro Franco andAranda Escobar 1984, 1985; Castro Franco and Bus-tos Zagal 1994, 2003; Davis and Dixon 1961; Davisand Smith 1953; Flores Villela 1993; Flores Villelaand Gerez 1988, 1994; Flores-Villela and Hernn-dez-Garca 2006; Flores Villela et al. 1991; Garca-Vazquez et al. 2006; Hernndez Garca 1989; Khler

and Heimes 2002; Lemos-Espinal et al. 1997; Sal-daa de la Riva and Prez Ramos 1987; Sites et al.1992; Smith 1934, 1939, 1984), karyology (Cole1978), parasites (Pelaez et al. 1948), phylogenet-ics and evolution (Flores-Villela et al. 2000; Harmonet al. 2003; Larsen and Tanner 1974, 1975; Schulteet al. 2003; Sites et al. 1992; Wiens 1993, 1998,1999; Wiens and Reeder 1997; Wills 1977), repro-duction (Davis and Dixon 1961: Fitch 1970; Guilletteet al. 1980; Mndez-de la Cruz et al. 1998; Saldaade la Riva and Prez Ramos 1987; Smith et al.2003).

REMARKS. Specimens of this species were

known but mis-identified as S. scalaris, S. graciosus,S. gratiosusor S. aeneusfrom at least 1874 to 1934.Some were referred to S. jalapaeas late as 1939. Allof these species are of about the same size, andhave somewhat similar patterns, but morphologicallyochoterenaeandjalapaeare much different from theothers.

The correct ending of this patronym, under the rulesof the first (1926), second (1961) and third (1964) edi-tions of the International Code of Zoological Nomen-clature was ochoterenai, although the name was orig-inally spelled ochoterenae. Therefore in Smith andTaylor (1950b) the supposedly correct spelling wasintroduced as ochoterenai. The former spelling wouldhave been proper if the name were considered to belatinized, rather than modern, but the 1926 Codeexplicitly excluded considerations of latinization (the1961 and 1964 Codes did not treat the subject). Thespelling ochoterenai followed the unexceptional rule(1926, 1961) or recommendation (1964) distinguish-ing patronyms from matronyms of modern names.

Not until the fourth (1999) edition of the Code ap-peared was it required under the Code to give aneponym an ending conformant with Latin grammar ifthe name were considered latinized (Art. 31.1.1). The1999 Code explicitly states that its provisions super-cede those of previous editions. Therefore the properending of this patronym depends on whether thename was originally considered latinized or not. Itobviously was, and therefore the name under the1999 Code should be spelled ochoterenae. This con-clusion is not destabilizing, inasmuch as the aiend-ing was used 17 times in the tallied literature, the aeending 21 times. The aeending has been used asrecently as 2002, and the aiending as recently as

1995. ETYMOLOGY. The name honors Dr. Isaac Och-terena, former Director of the Instituto de Biologa ofthe Universidad Nacional Autnoma de Mxico, in re-cognition of numerous courtesies extended to Taylorand Smith in their collecting trip of 1932.

ACKNOWLEDGMENTS. We are much indebted toJ. Simmons, E.N. Smith and G. Schneider for check-ing the identifications of certain specimens; to E. Pr-ez Ramos for help in plotting localities in Guerrero;and to the curators of AMNH, CU, FMNH, IPN, KU,LSUMZ, MCZ, MSB, MVZ, TCNC, TCWC, UCM,

UIMNH, UMMZ, USNM and UTA for information ontheir holdings of this species (acronyms from Levitonet al. 1985).

LITERATURE CITED

Aguilar Bentez, S. 1990. Dimensiones Ecolgicasdel Estado de Morelos. DR Centro Reg. Invest.Multidisciplinarias, Univ. Nac. Auton. Mex, Cuer-navaca, Morelos.

Ahl, E. 1934. ber eine Sammlung von Reptilien undAmphibien aus Mexiko. Zool. Anz. 106:184186.

Bell, E.L., H.M. Smith, and D. Chiszar. 2003. Anannotated list of the species-group names applied

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MAP. The circle indicates the type-locality, the dots other

occurrence records.


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to the lizard genus Sceloporus. Acta Zool. Mex.(n.s.) 90:103174.

Bojrquez-Tapia, L.A., I. Azuara, E. Ezcurra, and O.Flores-Villela. 1995. Identifying conservation pri-orities in Mexico through geographic informationsystems and modeling. Ecol. Appl. 5:215231.

Camarillo-Rangel, J.L. and R. Aguilar C. 1992. Note-

worthy distributional records for some Mexicanamphibians and reptiles. Southwest. Herpetol.Soc. Newsl. 22(1):56.

and H.M. Smith. 1992. A handlist of the amphibiansand reptiles of the state of Mexico, Mexico, p.3941. InP.D. and J.L. Strimple (eds.), Contribu-tions in Herpetology. Greater Cincinnati Herpetol.Soc., Cincinnati, Ohio.

Casas-Andreu, G. 1982. Anfibios y reptiles de la cos-ta suroeste del estado de Jalisco, con aspectossobre su ecologa y biogeografa. Doctoral Diss.UNAM, Mxico, D.F.

, X. Aguilar-Miguel and E.O. Pineda-Arredondo.1997. Lista taxonmica de los vertebrados terre-

stres del estado de Mxico. I. Anfibios y reptiles.Coleccin Ciencias y Tcnicas, Univ. Autn. Mx-ico, Toluca (32):953.

, F.R. Mndez-de la Cruz, and X. Aguilar-Miguel.2004. Anfibios y Reptiles, p. 375390. In A.J.Garca-Mendoza, M.d. Jess Ordez, y M.Briones-Salas (eds.), Biodiversidad de Oaxaca.Inst. Biol. UNAM, Fond. Oax Conserv Nat. WWF, Mxico.

Castro Franco, R. 2002. Historia natural de lagartijasdel estado de Morelos, Mexico. Unpubl. MastersThesis UNAM, Mexico.

and E. Aranda Escobar. 1984. Estudio preliminarsobre la ecologa de los reptiles del Estado de

Morelos. Unpubl. Bachelors Diss., Escuela deCiencias Biolgicas, Universidad Autnoma delEstado de Morelos, Cuernavaca.

and . 1985. Distribucin ecolgica de las lagartijasen el estado de Morelos. Mem. 8th Congr. Nac.Zool., Saltillo, Coahuila, Mxico:897908.

and M.G. Bustos Zagal. 1994. List of reptiles ofMorelos, Mexico, and their distribution in relationto vegetation types. Southwest. Nat. 39:171175.

and . 2003. Lagartijas de Morelos, Mxico: dis-tribucin, habitat y conservacin. Acta Zool. Mex.(n.s.) 88:123142.

, G. Garca Muoz, R. Lara Lpez, and G. BustosZagal. 1986. La herpetologa de Morelos: ecolo-ga e importancia econmica. Resmenes Prim-era Reunin Nat. Herpetol.:19.

Cochran, D.M. 1961. Type specimens of reptiles andamphibians in the U.S. National Museum. Bull.U.S. Natl. Mus. (220):xv + 291 p.

Cole, C.J. 1978. Karyotypes and systematics of thelizards in the variabilis, jalapae, and scalarisspe-cies groups of the genus Sceloporus. Amer. Mus.Novitates (2653):113.

Cope, E.D. 1887. Catalogue of batrachians and rep-tiles of Central America and Mexico. Bull. U.S.Natl. Mus. (32):198.

Cuesta Terron, C. 1932. Los camaleones mexicanos.An. Inst. Biol. Univ. Mexico 3:95121.

Davis, W.B. and J.R. Dixon. 1961. Reptiles (exclusiveof snakes) from the Chilpancingo region, Mexico.Proc. Biol. Soc. Washington 74:3756.

and H.M. Smith. 1953. Lizards and turtles of theMexican state of Morelos. Herpetologica 9:100108.

Duellman, W.E. 1965. Abiogeographic account of the

herpetofauna of Michoacn, Mexico. Univ. Kan-sas Publ. Mus. Nat. Hist. 15:627709.Dumril, A.H.A., M-F. Bocourt, and F. Mocquard.

18701909. tude sur les Reptiles. Mission Sci-entifique au Mexique et dans lAmrique Centrale.Recherches Zoologiques. Troisime Partie 1reSection. Impr. Nat., Paris.

Ferrari-Prez, F. 1886. Catalogue of animals collect-ed by the Geographical and Exploring Commis-sion of the Republic of Mexico. Proc. U.S. Natl.Mus. 9:125199.

Fitch, H.S. 1970. Reproductive cycles in lizards andsnakes. Univ. Kansas Publ. Mus. Nat. Hist. 52:1247.

Flores Villela, O.A. 1993. Herpetofauna Mexicana:annotated list of the species of amphibians andreptiles of Mexico, recent taxonomic changes,and new species. Carnegie Mus. Nat. Hist. Spec.Publ. (17):iv + 73 p.

and P. Grez. 1988. Conservacin en Mxico: sn-tesis sobre vertebrados terrestres, vegetacin y eluso del suelo. Inst. Nac. Investigaciones Recur-sos Nat., Xalapa, Mxico.

and . 1994. Biodiversidad y conservacin enMxico: vertebrados, vegetacin y uso del suelo.Univ. Nac. Autn. Mxico - UBIPRO, Mxico, D.F.

and E. Hernndez Garca. 2006. Herpetofauna dela Sierra de Taxco, Guerrero-Estado de Mxico,

p. 266282. InA. Ramrez-Bautista, L. Canseco-Mrquez, and F. Mendoza-Quijano (eds.), Inven-tarios herpetofaunsticos de Mxico: avances enel conocimiento de su biodiversidad. Publ. Soc.Herpetol. Mex. (3).

, , and A. Nieto-Montes de Oca. 1991. Catlogo deanfibios y reptiles del Museo de Zoologa, Facul-tad de Ciencias, Universidad Nacional Autnomade Mxico. Ser. Catlogos Mus. Zool. Alfonso L.Herrera (3):xii + 222 p.

, K.M. Kjer, M. Benabib, and J.W. Sites, Jr. 2000.Multiple data sets, congruence, and hypothesistesting for the phylogeny of basal groups of thelizard genus Sceloporus (Squamata, Phrynoso-matidae). Syst. Biol. 49:713739.

, F. Mendoza Quijano, and G. Gonzalez Porter(compl.). 1995. Recopilacion de claves para ladeterminacion de anfibios y reptiles de Mexico.Publ. esp. Mus. Zool. (10):iv + 285 p.

Frank, N. and E. Ramus. 1995. A Complete Guide toScientific and Common Names of Reptiles andAmphibians of the World. NG Publ., Inc., Potts-ville, Pennsylvania.

Gadow, H.F. 1905. The distribution of Mexican amph-ibians and reptiles. Proc. Zool. Soc. Lond. 1905:191245.

Garca, A. 2006. Using ecological niche modelling toidentify diversity hotspots for the herpetofauna of

855.3


4/5

Pacific lowlands and adjacent interior valleys ofMexico. Biol. Conserv. 130:25-46.

Garca-Vazquez, U.O., L. Canseco-Mrquez, J.L.Aguilar-Lpez, C.A. Hernndez-Jimnez, J. Ma-ceda-Cruz, M.G. Gutirrez-Mayn, and E.Y. Mel-garejo-Velez. 2006. Anlisis la distribucin de laherpetofauna en la regin Mixteca de Puebla,

Mxico, p. 152169. In A. Ramrez-Bautista, L.Canseco-Mrquez, and F. Mendoza-Quijano(eds.), Inventarios herpetofaunsticos de Mxico:avances en el conocimiento de su biodiversidad.Publ. Soc. Herpetol. Mex. (3).

Guillette, L.J., Jr., R.E. Jones, K.T. Fitzgerald, andH.M. Smith. 1980. Evolution of viviparity in thelizard genus Sceloporus. Herpetologica 36:201215.

Gnther, A.C.L.G. 1890. Biologia Centrali-Americana.Reptilia and Batrachia. R.H. Porter, and Dulauand Co., London.

Hernndez Garca, E. 1989. Herpetofauna de la Si-erra de Taxco, Gro. Prof. Diss., Biologist, Univ.

Nac. Aut. Mxico, Cd. Universitaria, D.F., Mxico.Harmon, L.J., J.A. Schulte, II, A. Larson, and J.B.Losos. 2003. Tempo and mode of evolutionary ra-diation in iguanian lizards. Science 301:961964.

Hutchins, M., J.B. Murphy, and N. Schlager (eds.).2003. Grzimeks Animal Life Encyclopedia. 2nded. Vol.7. Reptiles. Gale Group, Inc., FarmingtonHills, Michigan.

Khler, G. and P. Heimes. 2002. Stachelleguane:Lebensweise.Pflege.Zucht. Herpeton, Offenbach,Germany.

Larsen, K.R. and W.W. Tanner. 1974. Numeric analy-sis of the lizard genus Sceloporuswith special ref-erence to cranial osteology. Great Basin Nat. 34:

141. and . 1975. Evolution of sceloporine lizards(Iguanidae). Great Basin Nat. 35:120.

Lemos-Espinal, J.A., G.R. Smith, and R.E. Ballinger.1997. Thermal ecology of the lizard, Sceloporusgadoviae, in a tropical scrub forest. J. Arid Envi-ron. 35:311319.

Leviton, A.E., R.H. Gibbs, Jr., E. Heal, and C.E. Daw-son. 1985. Standards in herpetology and ichthyol-ogy: Part I. Standard symbolic codes for institu-tional resource collections in herpetology and ich-thyology. Copeia 1985:802821.

Liner, E.A. 1994. Scientific and common names forthe amphibians and reptiles of Mexico in Englishand Spanish. Nombres cientficos y comunes enIngls y Espaol de los anfibios y reptiles deMxico. SSAR Herpetol. Circ. (23):v + 113 p.

. 1996. Mexico bound IX. The Herper. LouisianaGulf Coast Herpetol. Soc. 3(4):712.

. 2007. A checklist of the amphibians and reptiles ofMexico. Occ. Pap. Mus. Nat. Sci. Louisiana St.Univ. (80):160.

Marx, H. 1976. Supplementary catalogue of type spe-cimens of reptiles and amphibians in Field Mus-eum of Natural History. Fieldiana Zool. 69:3394.

Mndez-de la Cruz, F.R., M. Villagrn-Santa Cruz,and R.M. Andrews. 1998. Evolution of viviparity inthe lizard genus Sceloporus. Herpetologica 54:

521532.Olson, R.E., B. Marx, and R. Rome. 1986. Descrip-

tive dentition morphology of lizards of Middle andNorth America, I: Scincidae, Teiidae, and Heloder-matidae. Bull. Maryland Herpetol. Soc. 22:97124

, , and . 1987. Descriptive dentition morphology oflizards of Middle and North America II: Igu-

anidae. Bull. Maryland Herpetol. Soc. 23:1234.Pelaez, D., R. Prez Reyes, and A. Barrera. 1948.Estudios sobre hematozoarios I. PlasmodiummexicanumThompson y Huff, 1944, en sus hus-pedes naturales. An. Esc. nac. Cienc. Biol. 5:197215.

Purdue, J.R. and C.C. Carpenter. 1972. A compara-tive study of the display motion in the iguanid gen-era Sceloporus, Uta, and Urosaurus. Herpetolo-gica 28:137141.

Saldaa de la Riva, L. and E. Prez Ramos. 1987.Herpetofauna del Estado de Guerrero, Mxico.Tesis Prof. Biol. UNAM, Mexico, D.F.

Schulte, J.A., II, J.P. Valladares, and A. Larson. 2003.

Phylogenetic relationships within Iguanidae infer-red using molecular and morphological data anda phylogenetic taxonomy of iguanian lizards.Herpetologica 59:399419.

Sites, J.W., Jr., J.W. Archie, C.J. Cole, and O. FloresVillela. 1992. A review of phylogenetic hypothesesfor lizards of the genus Sceloporus (Phrynoso-matidae): implications for ecological and evolu-tionary studies. Bull. Amer. Mus. Nat. Hist. (213):1110.

Smith, G.R., J.A. Lemos-Espinal, and R.E. Ballinger.2003. Body size, sexual dimorphism, and clutchsize in two populations of the lizard Sceloporusochoteranae. Southwest. Nat. 48:123126.

Smith, H.M. 1934. Descriptions of new lizards of gen-us Sceloporusfrom Mexico and southern UnitedStates. Trans. Kansas Acad. Sci. 37:263285.

. 1936. Descriptions of new species of lizards of thegenus Sceloporus from Mexico. Proc. Biol. Soc.Washington 49:8796.

. 1939. The Mexican and Central American lizards ofthe genus Sceloporus. Zool. Ser. Field Mus. Nat.Hist. (26): 1397.

. 1984. Notes on the enigmatic Barisia imbricataofthe British Museum, and on its collection of rep-tiles from Amula, Guerrero, Mexico. Bull. Mary-land Herpetol. Soc. 20:152158.

. 1987. Current nomenclature for the names andmaterial cited in Gnthers Reptilia and Batrachiavolume of the Biologia Centrali-Americana, p.XXIIILI. In A.C.L.G Gnther. 18851902. Bio-logia Centrali-Americana. SSAR Facs. ReprintsHerpetol., Ithaca, New York.

. 1991. The status of the southern Mexican lizardSceloporus carinatus. Bull. Maryland Herpetol.Soc. 27:195200.

, D.A. Langebartel, and K.L. Williams. 1964. Her-petological type-specimens in the University of Illi-nois Museum of Natural History. Illinois Biol.Monogr. (32):180.

, C. McCarthy, and D. Chiszar. 2000. Some enig-matic identifications in Boulenger's 1897 Scelo-

855.4


5/5

porus monograph (Reptilia: Sauria). Bull. Mary-land Herpetol. Soc. 36:124132.

and R.B. Smith. 1976. Synopsis of the Herpeto-fauna of Mexico. Source Analysis and Index forMexican Reptiles. Vol. III. John Johnson, NorthBennington, Vermont.

and . 1993. Synopsis of the Herpetofauna of Mexi-

co. Vol. VII. Bibliographic Addendum IV and In-dex, Bibliographic Addenda II-IV. Univ. Press ofColorado, Niwot, Colorado.

and E.H. Taylor. 1950a. Type localities of Mexicanreptiles and amphibians. Univ. Kansas Sci. Bull.33:313380.

and . 1950b. An annotated checklist and key to thereptiles of Mexico exclusive of the snakes. Bull. U.S. Natl. Mus. (199):v + 253 p.

Sokolov, V.E. (ed.). 1988. Dictionary of AnimalNames in Five Languages. Amphibians and Rep-tiles. Russky Yazyk Publ., Moscow.

Taylor, E.H. 1944. Present location of certain herpeto-logical and other types specimens. Univ. KansasSci. Bull. 30:117187.

Thomas, R.A. and J.R. Dixon. 1976. A reevaluationof the Sceloporus scalarisgroup (Sauria: Iguan-idae). Southwest. Nat. 20:523536.

Wellstead, C.F. 1982. Lizards from the Lower Valen-tine Formation (Miocene) of northern Nebraska. J.Herpetol. 16:364375.

Wiens, J.J. 1993. Phylogenetic relationships of phry-nosomatid lizards and monophyly of the Scelo-porusgroup. Copeia 1993:287299.

. 1998. Testing phylogenetic methods with tree con-gruence: phylogenetic analysis of polymorphicmorphological characters in phrynosomatid liz-

ards. Syst. Biol. 47:427444.. 1999. Phylogenetic evidence for multiple losses ofa sexually selected character in phrynosomatidlizards. Proc. R. Soc. Lond. B 266:15291535.

and T.W. Reeder. 1997. Phylogeny of the spiny liz-ards (Sceloporus) based on molecular and mor-phological evidence. Herpetol. Monogr. (11):1101.

Wills, F.H. 1977. Distribution, geographic variationand natural history of Sceloporus parvus Smith(Sauria: Iguanidae). M.S. Thesis, Texas A&M Uni-versity, College Station.

Oscar FloresVillela, Museo de Zoologa, Facultadde Ciencias, UNAM, A.P. 70399, Mxico, D.F. 04510Mxico ([email protected]); Hobart M.Smith, Department of EPO Biology, University ofColorado, Boulder, CO 803090334 ([email protected]); Ernest A. Liner, 310 Malibou Bou-levard, Houma, LA 703642598 ([email protected]); David Chiszar, Department of Psychology,University of Colorado, Boulder, CO 803090345([email protected]).

Primary editor for this account, Andrew H. Price.

Published 30 April 2008 and Copyright 2008 by the

Society for the Study of Amphibians and Reptiles.

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