METAMATERIALS

Quantum metasurface formultiphoton interferenceand state reconstructionKai Wang1, James G. Titchener1,2, Sergey S. Kruk1, Lei Xu1,3, Hung-Pin Chung1,4,Matthew Parry1, Ivan I. Kravchenko5, Yen-Hung Chen4,6, Alexander S. Solntsev1,7,Yuri S. Kivshar1, Dragomir N. Neshev1, Andrey A. Sukhorukov1*

Metasurfaces based on resonant nanophotonic structures have enabled innovative typesof flat-optics devices that often outperform the capabilities of bulk components, yet theseadvances remain largely unexplored for quantum applications. We show that nonclassicalmultiphoton interferences can be achieved at the subwavelength scale in all-dielectricmetasurfaces. We simultaneously image multiple projections of quantum states with asingle metasurface, enabling a robust reconstruction of amplitude, phase, coherence, andentanglement of multiphoton polarization-encoded states. One- and two-photon statesare reconstructed through nonlocal photon correlation measurements with polarization-insensitive click detectors positioned after the metasurface, and the scalability tohigher photon numbers is established theoretically. Our work illustrates the feasibilityof ultrathin quantum metadevices for the manipulation and measurement of multiphotonquantum states, with applications in free-space quantum imaging and communications.

The field of nanostructured metasurfacesoffers the possibility of replacing tradi-tionally bulky imaging systems with flatoptics devices (1), achieving high transmis-sion based on all-dielectric platforms (2–7).

Themetasurfaces provide a freedom to tailor thelight interference by coherently selecting andmix-ing different components on a subwavelengthscale, enabling polarization-spatial conversion(4, 7–12) and spin-orbital transformation (13).Such capabilitiesmotivatedmultiple applicationsfor the regime of classical light, yet the metasur-faces have the potential to emerge as essentialcomponents for quantum photonics (14–17).The key manifestations of quantum light are

associated with nonclassical multiphoton inter-ference, which is an enabling phenomenon for thetransformation and measurement of quantumstates. Conventionally, manipulation of multi-photon states is performed through a sequenceof beam-splitting optical elements, each realizingquantum interference (18–20). Recent advancesin nanotechnology have enabled the integration

of beam-splitters and couplers on tailored plas-monic structures (21, 22); however, materiallosses and complex photon-plasmon couplinginterfaces restrict the platform scalability. Werealize several multiphoton interferences in asingle flat all-dielectricmetasurface. The parallelquantum state transformations are encoded inmultiple interleaved metagratings, taking ad-vantage of the transverse spatial coherence ofthe photon wave functions extending across thebeam cross section. In the classical context, theinterleaving approach was effectively used forpolarization-sensitive beam splitting (8, 9, 11, 12),yet it requires nontrivial development for theapplication to multiphoton states.We formulate and realize an application of the

metasurface-based interferences formultiphotonquantum statemeasurement and reconstruction.We develop a metasurface that incorporates aset ofM/2 interleavedmetagratings [see part 3 of(23)], where M is an even number of diffractedbeams forming imaging spots. Each metasurfaceis composed of nanoresonators with speciallyvarying dimensions and orientations, accord-ing to the principle of geometric phase (8), tosplit specific elliptical polarization states (7),which would not be possible with conventionalgratings [see parts 1 and 2 of (23)]. This per-forms quantum projections in a multiphotonHilbert space to M imaging spots, which canbe considered as output ports. Each port cor-responds to a different elliptical polarizationstate (Fig. 1A), which is essential to minimize theerror amplification in quantum state recon-struction (24). Then, by directly measuring allpossible N-photon correlations, where N is thenumber of photons, from the M output beams,it becomes possible to reconstruct the initialN-photon density matrix, providing full infor-

mation on the multiphoton quantum entangle-ment. For example, in Fig. 1B, we show a sketchof three gratings (top) which realize an optimalset of projective bases shown as vectors on thePoincaré sphere (middle) forM = 6.The photon correlations betweenM output

ports can be obtained with simple polarization-insensitive single-photon click detectors. Theme-tasurface can be potentially combinedwith singlephoton–sensitive electron-multiplying charge-coupled device (EMCCD) cameras (25, 26) to de-termine the spatial correlations by processingmultiple time-frame images of quantum states.We consider quantum states with a fixed pho-ton numberN, which is a widely used approachin photon detection (27–30). The N-fold correla-tion data, stored in an array withN dimensions,are obtained by averaging the coincidenceevents over multiple time frames. For example,in Fig. 1C, we sketch a case with N = 2 photonsandM = 6. In each frame, two photons arrive atdifferent combinations of spots. After summingup the coincidence events over multiple timeframes,we obtain a correlation in two-dimensionalspace. Following the general measurement the-ory of (30), we establish that, for an indistin-guishable detection of N-photon polarizationstates (i.e., the detectors cannot distinguishwhich is which of the N photons), the requirednumber of output ports to perform the recon-struction scales linearly with the photon numberasM ≥ N + 3 (see Fig. 1B, bottom). For instance,with M = 6, up to N = 3 photon states can bemeasured.The parallel realization of multiphoton inter-

ferences with a single metasurface offers practicaladvantages for quantum state measurements.Conventional quantum state tomography (27)methods based on reconfigurable setups canrequire extra time and potentially suffer fromerrors associated with the movement of bulkoptical components (27) or tuning of optical in-terference elements (31). Moreover, the conven-tionally used sequential implementations ofprojective measurements present a fundamen-tal limit for miniaturization while being inher-ently sensitive to fluctuations or misalignmentbetween different elements, especially for higher–photon number states. The emerging methodsbased on static transformations implementedwith bulk optical components (19) or integratedwaveguides (28–30) still require multiple stagesof interferences. By contrast, our quantum meta-surface provides an ultimately robust and com-pact solution, the speed of which is limited onlyby the detectors.We fabricate silicon-on-glass metasurfaces de-

signed forM = 6 and 8 imaging spots using stan-dard semiconductor fabrication technology [seeparts 4 and 7 of (23) for details]. The experimentallydetermined polarization projective bases obtainedthrough classical characterization are plotted onthe Poincaré sphere in Fig. 2A for a metasurfacewith M = 6 that is used later for quantum ex-periments. The transfer matrix measurementsconfirm that the polarization projective basesare close to the optimal frame. The condition

RESEARCH

Wang et al., Science 361, 1104–1108 (2018) 14 September 2018 1 of 4

1Nonlinear Physics Centre, Research School of Physics andEngineering, The Australian National University, Canberra,ACT 2601, Australia. 2Quantum Technology EnterpriseCentre, Quantum Engineering Technology Labs, H. H. WillsPhysics Laboratory and Department of Electrical andElectronic Engineering, University of Bristol, Bristol BS8 1FD,UK. 3School of Engineering and Information Technology,University of New South Wales, Canberra, ACT 2600,Australia. 4Department of Optics and Photonics, NationalCentral University, Jhongli 320, Taiwan. 5Center forNanophase Materials Sciences, Oak Ridge NationalLaboratory, Oak Ridge, TN 37831, USA. 6Center forAstronautical Physics and Engineering, National CentralUniversity, Jhongli 320, Taiwan. 7School of Mathematicaland Physical Sciences, University of Technology Sydney,Ultimo, NSW 2007, Australia.*Corresponding author. Email: andrey.sukhorukov@anu.edu.au

on October 10, 2020

http://science.sciencem

ag.org/D

ownloaded from

number, a measure of error amplification inthe reconstruction [see part 1 of (23)], is 2.08,close to the fundamental theoretical minimumof

ffiffiffi

3p

≃ 1:73. The reconstruction is immune tofabrication imperfections because their effect isfully taken into consideration by performing anexperimental metasurface characterization withclassical light after the fabrication [see parts 6and 10 of (23)].

First, we show that our metasurface enables ac-curate reconstruction of the quantum-polarizationstate of single photons. A heralded photon sourceis used at awavelength of 1570.6 nmon the basis ofspontaneous parametric down conversion (SPDC)in a nonlinear waveguide [see parts 5, 8, 9, and 11of (23) for details]. The heralded single photonsare initially linearly polarized. They are preparedin different polarization states by varying the

angle of a quarter-wave plate (QWP), then sentto the metasurface, and each diffracted photonbeam is collected by a fiber-coupled interfaceto the single-photon detectors. Bymeasuring thecorrelations with the master detector, we recon-struct the quantum-polarization state from thephoton counts at the six ports. The results areshown in Fig. 2B, where the curves are theo-retical predictions and dots are experimental

Wang et al., Science 361, 1104–1108 (2018) 14 September 2018 2 of 4

Fig. 2. Experimental measurement ofheralded single-photon states with themetasurface. (A) Classically characterizedprojective bases of the metasurface forports numbered 1 to 6. (B) Accumulatedsingle-photon counts in each of M = 6output ports versus the angle of a QWPrealizing a photon state transformationbefore the metasurface. Experimentaldata are shown as dots, with error barsindicating shot noise. Solid lines representtheoretical predictions based on classicallymeasured metasurface transfer matrix.rad, radians. (C) Comparison between theprepared (solid line) and reconstructed(dots) states based on the measurementspresented in (B), plotted on a Poincaré sphere.

A

Average fidelity 99.35%(96.44% to 99.98%)

Condition number 2.08

B

Port 1

Port 6

Port 2

Port 5

Port 4

Port 3

C

1

6

52

3

4

6

3

B C

1 2 3 4 5 6

1

2

3

4

5

6

Correlation

1 2 3 4 5 6

Frame 1

Frame 3

Frame 4

Frame 2

A

N-photon polarization state

Reconstruction

Metasurface

N-fold correlation

12

M

Density matrix

ρ

1 µm

Fig. 1. Concept of quantum state imaging via nanostructured flatoptics. (A) Sketch of a metasurface being used to image an input N-photonpolarization state into anM-spot image. At the top right is a scanning electronmicroscopy image of the fabricated all-dielectric metasurface. Greencrosses represent photons; purple blocks on the metasurface representnanoresonators. (B) The top image is a sketch of three interleaved gratingsfor M = 6.The middle image shows, with orange arrows, the corresponding

projective bases as vectors on the Poincaré sphere; black arrows indicatethe coordinate axes. Shown at the bottom are the minimum number ofrequired spots to fully reconstruct the initial quantum state for differentnumbers of photons N, where optimal-frame choice of projective basesexists for M = 6, 8, 12, 20, … . (C) An example of correlation measurementwith N = 2 andM = 6, with several time-frame measurements combinedinto a two-dimensional correlation image.

RESEARCH | REPORTon O

ctober 10, 2020

http://science.sciencemag.org/

Dow

nloaded from

measurements. We observe that the measure-ment errors are dominated by the single-photondetection shot noise, which is proportional to thesquare root of the photon counts, as indicated bythe error bars. We use the measured photoncounts to reconstruct the input single-photonstates by performing a maximum-likelihood es-timation (27) and plot them on a Poincaré spherein Fig. 2C. The reconstructed states present a

high average fidelity of 99.35% with respect tothe prepared states.Next, we realize two-photon interference, the

setup ofwhich is conceptually sketched in Fig. 3A.The SPDC source generates a photon pair withhorizontal (H) and vertical (V) polarizations, withthe path length difference between polarizationcomponents controllable by a delay line [see part12 of (23) for details]. We measure the effect of

delay on the two-photon interference, analogousto the Hong-Ou-Mandel (HOM) experiment (32).In such a nontrivially generalized two-photoninterference, we expect a dip or peak dependingon the two-by-two transfer matrixTa;bº½ua;ub�†from the two-dimensional polarization statevector to a chosen pair of ports, where † denotestranspose conjugate and ua and ub are the pro-jective bases of ports a and b, respectively. We

Wang et al., Science 361, 1104–1108 (2018) 14 September 2018 3 of 4

A

B C

Port 1&6 Port 1&5

Fidelity 95.24%

EDRe(ρ), Measured Im(ρ), Measured

Im(ρ), Predicted

Fidelity 98.54%

GFRe(ρ), Measured

Re(ρ), Predicted

Im(ρ), Measured

Im(ρ), Predicted

Re(ρ), Predicted

SPDC waveguide

Pump filtering

Photon pair generation

Delay-line State measurementSState measumeasuremeementState measurementTransformation

Pumplaser

Mirror

PBS

Prism

Mirror

Mirror

Prism on a linear stage

PBS

MetasurfaceFiber couplers

APD

APD

V

H

12

3

45

6

QWP

1 2 3 4 5

2

3

4

5

6

Port number

rebmun troP

0

217

Counts

0

235

Counts1 2 3 4 5

2

3

4

5

6

Port number

rebmun troP

Fig. 3. Experimental two-photon interferences and state reconstructionwith the metasurface. (A) Schematic of the setup, including photon-pairgeneration and pump filtering, a delay line with polarizing beam splitters(PBSs) to control the path difference between orthogonally polarized photonsin a pair, state transformation with a QWP, and state measurement withthe metasurface using avalanche photodiodes (APDs). (B and C) Quantumcorrelations between ports 1 and 6 (B) with close-to-orthogonal bases andports 1 and 5 (C) with nonorthogonal bases, shown with dots and error bars

indicating shot noise. Solid curves represent theoretical predictions. Redarrows in the Poincaré spheres denote projective bases of different ports.Blue arrows indicate the polarization state of entangled photons, withone photon in H and the other in V polarization. (D and F) Representativetwofold correlation measurements and (E and G) the correspondingreconstructed density matrices r labeled “Measured” alongside thetheoretically predicted states labeled “Predicted” for QWP orientationsq = 0° [(D) and (E)] and q = 37.5° [(F) and (G)].

RESEARCH | REPORTon O

ctober 10, 2020

http://science.sciencemag.org/

Dow

nloaded from

note that Ta,b corresponds to an effective Hermi-tian Hamiltonian only if ua and ub are orthogo-nal, resulting in a conventionalHOMdip,whereasotherwise, a HOMpeak can appear analogous toa lossy beam splitter (22). Here we set the angleof the QWP at q = 0°, which means that thephoton pairs are in a state r(q = 0°), where onephoton isHpolarized and another is V polarized.As reflected in the Poincaré sphere of Fig. 3B—where the red arrows denote projective bases ofthe two ports (ua and ub) and the blue arrowsrepresent the polarization of the photon pairs,one photon inH and the other in Vpolarization—we see that the state vector u1 points in the oppo-site direction of u6. We find that, in this case,photons with cross-polarized entanglement inH-V basis will give rise to a dip in the interfer-ence pattern with the variation of path-lengthdifference (see Fig. 3B, left). Such a behavioris directly caused by the coalescence nature ofbosons. The situation is quite different if wemeasure such an interference between portsa = 1 and b = 5, because u1 and u5 are far frombeing orthogonal. This can be seen from the redarrows in the Poincaré sphere of Fig. 3C, wherethe angle between the two vectors representingu1 and u5 is much smaller than p. For entangledphotons with H and V polarization in a pair, in-terference under the transfer matrix T1,5 leads toa peak instead of a dip when varying the pathdifference in the delay line. Indeed, in Fig. 3C(left) we observe a peak, which is related to theanticoalescence of bosons in transformationsinduced by non-HermitianHamiltonians, a non-trivial generalization of the HOM interferenceanalogous to (22). For details of the theoreticalpredictions and experimentalmethods, see part 4of (23).As a following step, wemeasure all 15 twofold

nonlocal correlations between theM = 6 outputsfrom themetasurface for a given input statewherethe time delay is fixed to zero. This provides us fullinformation to accurately reconstruct the inputtwo-photon density matrix. We use two single-photon detectors tomap out all possible outputcombinations, although this could potentially beaccomplished evenmore easily with an EMCCDcamera. We show representative results for twodifferent states r(q = 0°) and r(q = 37.5°) in Fig. 3,D and E, and Fig. 3, F and G, respectively. Notethat r(q = 0°) is a state in which photon pairshave cross-polarized entanglement beyond theclassical limit, yet it is not fully pure [see part 4of (23)], providing a suitable test case for recon-struction of general mixed states. In Fig. 3D, weshow the measured twofold correlations for theinput state r(q = 0°), and the reconstructed den-sity matrix is shown in Fig. 3E. That only the

bunched four central elements are nonzero con-firms the cross-polarized property of our pho-ton pairs in H-V basis. Moreover, the nonzerojHVVHi element implies the presence of two-photon entanglement. It is smaller compared tothe diagonal element jHVHVi , indicating thatthe polarization state is not fully pure. Althoughr(q = 0°) only has nonzero elements in the realpart of the density matrix, we also show themeasurement and reconstruction of r(q = 37.5°),which contains nontrivial imaginary elements,in Fig. 3, F and G. In both cases, we achieve avery good agreement between the predicted andreconstructed density matrices, as evidencedby high fidelity exceeding 95%. The correlationcounts are obtained by a Gaussian fitting to thecorrelation histogram to remove the background,which is less than 10% of the signal for all mea-surements shown in Fig. 3F; see details in part 12of (23).Our results illustrate themanifestation ofmulti-

photon quantum interference onmetasurfaces.We formulate a concept of parallel quantumstate transformation with metasurfaces, enablingsingle- and multiphoton state measurementssolely based on the interaction of light with sub-wavelength thin nanostructures and nonlocal cor-relationmeasurements without a requirement ofphoton number–resolvable detectors. This pro-vides ultimateminiaturization and stability com-bined with high accuracy and robustness, as wedemonstrate experimentally via reconstruction ofone- and two-photonquantum-polarization states,including the amplitude, phase, coherence, andquantumentanglement. In general, our approachis particularly suitable for imaging-based mea-surements of multiphoton polarization states,where themetasurface can act as a quantum lensto transform the photons to a suitable format forthe camera to recognize and retrieve more in-formation. Furthermore, there is the potential tocapture other degrees of freedom associatedwithspatially varying polarization states for themanip-ulation and measurement of high-dimensionalquantum states of light, with applications in-cluding free-space communications and quan-tum imaging.

REFERENCES AND NOTES

1. N. Yu, F. Capasso, Nat. Mater. 13, 139–150 (2014).2. A. I. Kuznetsov, A. E. Miroshnichenko, M. L. Brongersma,

Y. S. Kivshar, B. Luk’yanchuk, Science 354, aag2472 (2016).3. M. Decker et al., Adv. Opt. Mater. 3, 813–820 (2015).4. A. Arbabi, Y. Horie, M. Bagheri, A. Faraon, Nat. Nanotechnol. 10,

937–943 (2015).5. S. Kruk et al., APL Photonics 1, 030801 (2016).6. P. Genevet, F. Capasso, F. Aieta, M. Khorasaninejad, R. Devlin,

Optica 4, 139 (2017).7. J. P. Balthasar Mueller, N. A. Rubin, R. C. Devlin, B. Groever,

F. Capasso, Phys. Rev. Lett. 118, 113901 (2017).

8. Z. Bomzon, V. Kleiner, E. Hasman, Opt. Lett. 26, 1424–1426(2001).

9. A. Pors, M. G. Nielsen, S. I. Bozhevolnyi, Optica 2, 716 (2015).10. J. P. Balthasar Mueller, K. Leosson, F. Capasso, Optica 3, 42

(2016).11. E. Maguid et al., Science 352, 1202–1206 (2016).12. F. Ding, A. Pors, Y. T. Chen, V. A. Zenin, S. I. Bozhevolnyi, ACS

Photonics 4, 943–949 (2017).13. R. C. Devlin, A. Ambrosio, N. A. Rubin, J. P. B. Mueller,

F. Capasso, Science 358, 896–901 (2017).14. P. K. Jha, X. Ni, C. Wu, Y. Wang, X. Zhang, Phys. Rev. Lett. 115,

025501 (2015).15. T. Roger et al., Nat. Commun. 6, 7031 (2015).16. A. Lyons et al., arXiv:1709.03428 [quant.-ph] (11 September

2017).17. T. Stav et al., Science 361, 1101–1104 (2018).18. J. W. Silverstone et al., Nat. Photonics 8, 104–108 (2014).19. O. Bayraktar, M. Swillo, C. Canalias, G. Bjork, Phys. Rev. A 94,

020105 (2016).20. J. S. Fakonas, A. Mitskovets, H. A. Atwater, New J. Phys. 17,

023002 (2015).21. G. Di Martino et al., Phys. Rev. Appl. 1, 034004 (2014).22. B. Vest et al., Science 356, 1373–1376 (2017).23. See supplementary materials.24. M. R. Foreman, A. Favaro, A. Aiello, Phys. Rev. Lett. 115,

263901 (2015).25. M. P. Edgar et al., Nat. Commun. 3, 984 (2012).26. M. Reichert, H. Defienne, J. W. Fleischer, Sci. Rep. 8, 7925

(2018).27. D. F. V. James, P. G. Kwiat, W. J. Munro, A. G. White, Phys. Rev. A

64, 052312 (2001).28. J. G. Titchener, A. S. Solntsev, A. A. Sukhorukov, Opt. Lett. 41,

4079–4082 (2016).29. D. Oren, M. Mutzafi, Y. C. Eldar, M. Segev, Optica 4, 993

(2017).30. J. G. Titchener et al., npj Quantum Inf. 4, 19 (2018).31. P. J. Shadbolt et al., Nat. Photonics 6, 45–49 (2012).32. C. K. Hong, Z. Y. Ou, L. Mandel, Phys. Rev. Lett. 59,

2044–2046 (1987).

ACKNOWLEDGMENTS

We gratefully thank H. Bachor, M. Scully, I. Walmsley, and F. Setzpfandtfor fruitful discussions; R. Schiek and Y. Zarate for help in developingovens for waveguide temperature control; and M. Liu for adviceon numerical simulations. Funding: This work was supported by theAustralian Research Council (including projects DP160100619,DP150103733, and DE180100070) and the Ministry of Scienceand Technology (MOST), Taiwan, under contract 106-2221-E-008-068-MY3. A portion of this research was conducted at the Center forNanophase Materials Sciences, which is a U.S. DOE Office of ScienceUser Facility. Author contributions: K.W., D.N.N., and A.A.S.conceived and designed the research; K.W. and L.X. performednumerical modeling of metasurface design; S.S.K. and I.I.K. fabricatedthe dielectric metasurfaces; H.-P.C. and Y.-H.C. fabricated nonlinearwaveguides; K.W., J.G.T., H.-P.C., M.P., and A.S.S. performed opticalexperimental measurements and data analysis; A.A.S., D.N.N., andY.S.K. supervised the work; K.W., A.A.S., D.N.N., and Y.S.K. prepared themanuscript and supplementary materials in coordination with allauthors. Competing interests: The authors declare no competinginterests. Data and materials availability: All data needed toevaluate the conclusions in this study are presented in the paperor in the supplementary materials.

SUPPLEMENTARY MATERIALS

www.sciencemag.org/content/361/6407/1104/suppl/DC1Materials and MethodsFigs. S1 to S11References (33–37)

10 April 2018; accepted 17 July 201810.1126/science.aat8196

Wang et al., Science 361, 1104–1108 (2018) 14 September 2018 4 of 4

RESEARCH | REPORTon O

ctober 10, 2020

http://science.sciencemag.org/

Dow

nloaded from

Quantum metasurface for multiphoton interference and state reconstruction

Alexander S. Solntsev, Yuri S. Kivshar, Dragomir N. Neshev and Andrey A. SukhorukovKai Wang, James G. Titchener, Sergey S. Kruk, Lei Xu, Hung-Pin Chung, Matthew Parry, Ivan I. Kravchenko, Yen-Hung Chen,

DOI: 10.1126/science.aat8196 (6407), 1104-1108.361Science 

, this issue p. 1104, p. 1101Scienceplatform.photons. The results should aid the development of integrated quantum optic circuits operating on a nanophotonic

used a dielectric metasurface to generate entanglement between spin and orbital angular momentum of singleet al.Stav multiple photons by simply passing them through a dielectric metasurface, scattering them into single-photon detectors.

determined the quantum state ofet al.that metasurfaces can be extended into the quantum optical regime. Wang Metasurfaces should allow wafer-thin surfaces to replace bulk optical components. Two reports now demonstrate

Going quantum with metamaterials

ARTICLE TOOLS http://science.sciencemag.org/content/361/6407/1104

MATERIALSSUPPLEMENTARY http://science.sciencemag.org/content/suppl/2018/09/12/361.6407.1104.DC1

CONTENTRELATED http://science.sciencemag.org/content/sci/361/6407/1101.full

REFERENCES

http://science.sciencemag.org/content/361/6407/1104#BIBLThis article cites 35 articles, 5 of which you can access for free

PERMISSIONS http://www.sciencemag.org/help/reprints-and-permissions

Terms of ServiceUse of this article is subject to the

is a registered trademark of AAAS.ScienceScience, 1200 New York Avenue NW, Washington, DC 20005. The title (print ISSN 0036-8075; online ISSN 1095-9203) is published by the American Association for the Advancement ofScience

Science. No claim to original U.S. Government WorksCopyright © 2018 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of

on October 10, 2020

http://science.sciencem

ag.org/D

ownloaded from