physiology of vision

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  • 2

    ()

  • 70%

    , , ,

  • Figure 15.6

  • 7

    6 (4 , 2 )

  • 2

    3

  • ,

    , ,

    ( , ) : + ()

  • 2 : ()

    ( 7 ) ( ,

    )

    ( )

  • -

  • 2

    (D) D=(n2-n1)/r n1: 1 n1: 2 r: m

  • ~ 60 D, , 24 mm

  • Figure 15.17a

    ,

    ~>7 m

  • :

    , ,

  • E ,

    M

  • Problems of Refraction

    Figure 15.18

  • ,

  • (

    )

    ( )

  • 3 3

  • Light

    Figure 15.14

  • (~16:1)

    : = +

  • > 100x106 1x106

  • -

  • :

    H

    : : 1:1:1

  • (Phototransduction)

    O ( )

  • Baylor et al, >106 +

  • 1 ?

  • ( A)

    : 11-cis and all-trans

  • Chemistry of Visual Pigments

    Figure 15.20

    , . cis-

    ,

  • (o + 11-cis )

    all-trans +

    ( , bleaching of the pigment)

    ll-trans 11-cis

    11-cis A

    11-cis + =>

  • Figure 15.21

  • ( + o)

    3 : blue, green, and red ( )

    3

  • all-trans ,

    G (transducin)

    (PDE)

    PDE cGMP GMP +

    cGMP, , ,

  • Phototransduction

    Figure 15.22

  • (d-30000 molecules/m2)

    ~109

    +

    G

    (11-cis-form)

  • ( ) :

  • (~80% )

    : ( , )

  • 2 => 2 ( ) =>

    2

    , ( )

  • On-center

    Off-center -

    on off

    :

  • Figure 15.24

    :