3

HJカKh비( A

HNo©

Register of Hosta Species and their Nomenclature • Part

For complete information refer to Species Update on the home page of Hostalibrary.org © 2008 • The American Hosta Society/W.G. Schmid

ower leaves.

. nakaiana F. Maekawa . Japanese Bot., 11:687–689; 1935 ンザシギボウシ = 簪擬宝珠 = anzashi Gibōshi = Ornamental airpin hosta. 녀비비추 = Bi-nyeo-bi-bi-chu

Korean)

HS name: H. nakaiana

. nakaiana (Cultivated) ► otice large, viable fertile bracts; pening buds W.G. Schmid; Hosta Hill 1991.

H. nakaiana speciated in the southern areas of the Japanese archipelago, when displaced specimens of H. capitata from Korea intergraded with Japanese species. The lack of papillose veins projecting the leaf underside points to that. H. nakaiana specimens now grown have smooth veins and are fertile so establish in the wild in disjunct Japanese areas. Several forms have evolved but they differ, somewith larger, others with narr

■ H. nigrescens Maekawa. J. of the Faculty of Science, Sect. 3 Botany, Vol. 5:352 -355, 1940. ◄ H. nigrescens (cultivated) Single bloom scape on young plant; Hosta Hill R.G. HH No. 12388604 ▪ © W.G. Schmid; 10 Aug. 1988.

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Japanese names: クロギボウシ = 黒擬宝珠 = Kuro Gibōshi = black hosta テンリュギボウシ = 天竜擬宝珠 = Tenryu Gibōshi ちょだいぎんいろギボウシ

= 著大銀色 = Chodai Giniro Gibōshi = H. ‘Tenryu’ AHS Name: H. nigrescens (also as H. ‘Nigrescens’)

H. nigrescens (cultivated) Hab. in mountain valleys in Glaucous leaf coating as seen in spring Tōhoku and Chūbu region Hosta Hill R.G. ▪ Spec. HH 05138205 (Chūbu-chihō; 中部地方), © W.G. Schmid Photo: 16 May 1987 the central region of Honshū (K10/20 in KYO; coll. in Aug 1903 by N. Kinashi); as H. ‘Tenryu’ in Aichi-ken (愛知県) and Shizuoka-ken (静岡県) and in Tenryu River area (天竜川). Sometimes considered a cultivar, but native populations have been observed in Tenryugawa and further north in Iwate and Aomori prefectures, all in the northern area of Honshū, Tōhoku-chihō (東北地方).

♠ H. nigrescens f. elatior Maekawa. J. of the Faculty of Science, Sect. 3 Botany, Vol. 5:355. さいたか クロギボウシ = 最多佳黒

擬宝珠 = Saitaka Kuro Gibōshi (Maek. 1940)

AHS name: H. nigrescens ‘Elatior’ Cultivar name: H. ‘Elatior’

H. ‘Elatior’ ► Early anthesis; scapes = 5 ft (1.5 m) Hosta Hill R.G. HH 01978504 (Kaneko 1971) 8 July 1987 ▪ © W.G. Schmid Photo

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Hab.: This culton was found by Mataroku Yamanaka in cultivation. Maekawa (1940) published this taxon name based on a cultivated plant (without number) in Kikuchi's garden in Kyoto. Representative wild populations have not been found. Morphometric analysis indicates this to be an interspecific hybrid between H. montana and H. nigrescens combining the long-scape trait of H. nigrescens with the flower and leaf characteristics and blooming time of H. montana. Schmid (1991) reduced this taxon to cultivar rank. At Hosta Hill R.G. (located at 1188 feet (361m) AMSL; 84-12'-30" W and 33-51' N), this cultivar blooms in late June to early July, a later than H. montana. Specimens of H. nigrescens do not bloom until early August.

♠ H. okamotoi Araki. Acta Phytotaxonomica et Geobotanica, Vol. 11, 321; 1942. H. okamotoi = オクヤマギボウシ = Okuyama Gibōshi = Mount Okuyama Hosta AHS Name: H. okamotoi Hab.: Province of H. okamotoi (BGO Voucher - Cultivated) Tamba (丹波国; Hosta Hill R.G. ● W.G. Schmid May 1991 Tamba-no kuni). Collected by Okamoto in the vicinity of the village of Chii-mura in Kitakuwada-gun (北桑田郡) in Kinki Region (近畿地方). Closely related to H. takiensis and H. aequinoctiiantha and with these encompasses the “late-blooming” component of section Picnolepis. It also has a similar habitat, adapted to thin soil on arocky substrate in forested, isomountain valleys on slopes, also grows epiphytically on tree trunks. Differentiated from the above taxa by RAPD/DNA analysis

lated

H. okamotoi (Cultivated) ► BGO Voucher Hosta Hill R.G. © 2004 H. Philips

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H. pachyscapa Maekawa. J. of the Faculty of Science, Imperial University Tokyo, Section 3 Botany, Vol. 5:383 ic. 58–59 1940. ベンケイギボウシ = 弁慶

擬宝珠 = Benkei Gibōshi = “Strong man hosta” (alluding to the thick scapes; pachy = thick; scapa = scapes). AHS Name: H. ‘Pachyscapa’ H. ‘Pachyscapa’ ► Detail of thick scape Summers No. 184 1969 at Hosta Hill R.G. © Photo W.G. Schmid June 1988 H. ‘Pachyscapa’ at Hosta Hill R.G. 2006 Schmid (2006) considers H. ‘Pachyscapa’ to be an interspecific hybrid population, which grows in limited areas of the Kinki region. Very low fertility (Zonneveld 2001) points to cultivated, hybrid origin and wild populations have not been confirmed by Fujita, other than being mentioned as synonyms and no further field data have been published since 1940. For this reason it is perhaps best to consider this to be a culton and it was classified as a culton in Schmid (1991), using the name as H. ‘Pachyscapa’.

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H. plantaginea (Lamarck) Ascherson Botanische Zeitung, 21:53, ic. Pl. 24 1863.

52).

chi 1934)

l. y

Type: Ill. in Ascherson, Pl. 24, 1863 (syn.); ITIS Taxon Serial No.: 429 AHS Name: H. plantaginea 玉簪 = ｷﾞｮｸｼﾝ = yu zan (Chinese)

HTypical Dist

Cou

棒玉簪 = ﾎﾞｳｷﾞｮｸｼﾝ = bang yu zan (Chinese Alias/Synonym) 玉春棒 = ｷﾞｮｸｼｭﾝﾎﾞｳ = yuchun bang (Chinese Alias/Synonym) 白鶴花 = ﾊｸｶｸｶ = bai he hua (Chinese Alias/Synonym) マルバタマノカンザシ = maruba tama no kanzashi (Japanese; Kiku옥잠화 = Ok-jam-hwa (Korean) 둥근옥잠화 = Dung-geun-ok-jam-hwa (Korean synonym) 비녀옥잠화 = Bi-nyeo-ok-jam-hwa (Korean synonym) Hab. and History: A native to China and cultivated since the Han Dynasty (漢朝 ▪ 206BC-220AD). The species was once thought to have evolved in Hebei province (河北), surrounding Beijing (北京; Běijīng = northerncapital), but Flora of China (Vo24, 2000) states: “Forests, grassslopes, rocky places; sea level to 2200 m. Some areGuang-dong (广东; eastern Guang); Guangxi (广西

)(?); Hubei (湖北; north of lake); Hunan (湖南; soLiaoning (遼寧)(?); Shaanxi (陕西(?); Qin Ling M(云南) (?); Zhejiang (浙江)(?). It is the first hosta grown from seeds sent from Macao by the French 1784. Korean native populations are not reported

FloH. pl

H. tibae flow(Left M

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. plantaginea (Cult.) ended Bud Stage; Developing Budsrt.: © Quingdao Nurs. 山東

青岛平度苗圃

an

목록 2004).

?. Anhui (安徽); Fujian (福建); ; western Guang); Hebei (河北

uth of lake); Jiangsu (江苏); ountains; Sichuan (四川); Yunnknown on the continent of Europe Consul Charles de Guignes before (KPNI - 국가표준식물

wer Size Comparison antaginea Flower in Center er left ▬ H. montana flower right

easure is 1 inch, Right is 1 cm)

H. plantaginea var. japonica Kikuchi ex Maekawa. J. of the Faculty of Science, Imperial University Tokyo, Section 3 Botany, Vol. 5:343, 344, 348; ic. 6, 12; 1940. タマノカンザシ = tama no kanzashi (Japanese; Ranzan Ono; 小野蘭山) 긴옥잠화 = Gin-ok-jam-hwa (Korean Synonym) AHS Name: H. plantaginea var. japonica In the late 1820s, von Siebold obtained a sample of H. plantaginea var. japonica while at Deshima (出島 = protruding island). In 1830, he deposited this plant at Batavia's Buitenzorg gardens where it was planted (Schmid 1991). In 1841 J. Pierot of the Rijksherbarium Leiden (L) retrieved H. plantaginea var. japonica andLeiden under the name Funkia grandiflora. This “Japanese Plantaginea” differs from the Chinese species by having longer and narrower leaf blaand making a taller, a less compact leaf mound; by the anthers being yellow with slight purple dotting; and by the lobes of H. plantaginea var. japonica, more narrow and straigh

H. plantaginea (cult.) Detail of Typical Floral Organs Anthers with Stamens and Pistil

sent it to

des

t looking and less ecurving (see photo page 7).

awa ce, Sect. 3

d

aru옥잠화 = Gin-ok-jam-hwa (Korean name).

HS Name: H. plantaginea ‘Stenantha’ (reduced t

morphological differentiation stenos = narrow and

r H. plantaginea f. stenantha Maek1940 J. of the Fac. of ScienBotany, Vol. 5:347, ic. 9 = H. plantaginea ‘Stenantha’ Schmi1991: The genus Hosta: Gibōshi Zoku (ギボウシ属); Timber Press pp. 85, 314. クダザキマルバタマノカンザシ = kudazaki m

Note

긴 A This phenotype is a natural modification of the H.established by Maekawa (1940). The species epithe

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H. plantaginea f. stenantha Lin rt Style

Court.: ©iroha.s7.xrea ear-Oblong Tepals-Sho

ba tama no kanzashi (Japanese).

o culton Schmid 1991)

to its minor anthos = flower.

plantaginea var. japonica. t stenantha points

H. plantaginea var. japonica Typical Slender Inflorescences and Flowers

Court.: © Tomobee’s PL

■H. pulchella Fujita. H. pulchella Fujita. Acta Phytotaxonomica et Geobotanica, Vol. 27, 3/4:89, ic. 8 1976.

Acta Phytotaxonomica et Geobotanica, Vol. 27, 3/4:89, ic. 8 1976. H. pulchella = ウバタケギボウシ

= 姥岳擬宝珠 = Ubatake Gibōshi H. pulchella = ウバタケギボウシ

= 姥岳擬宝珠 = Ubatake Gibōshi H. pulchella ‘Sobo’ hort. = 祖母姫 H. pulchella ‘Sobo’ hort. = 祖母姫

岩擬宝珠 = 祖母姫岩ギボウシ = 岩擬宝珠 = 祖母姫岩ギボウシ = ソボヒメイワギボウシ = Sobo Hime Iwa Gibōshi ソボヒメイワギボウシ = Sobo Hime Iwa Gibōshi AHS Name: H. pulchella AHS Name: H. pulchella Hab.: On mountain ridges and between rocks on thin soils over rock strata at 1600 m ASML. Allopatric in Kumamoto-ken

Hab.: On mountain ridges and between rocks on thin soils over rock strata at 1600 m ASML. Allopatric in Kumamoto-ken

H. pulchella (cultivated) UGA Voucher KYO (Currie No. 100) Coll. Sobosan (祖母山) Miyazaki-ken (宮崎県)

Hosta Hill R.G. ▪ © W.G. Schmid 5 May 1989

(熊本県), Miyazaki-ken(宮崎県) and in Ōita-ken (大分県) in western Kyūshū., Japan.

(熊本県), Miyazaki-ken(宮崎県) and in Ōita-ken (大分県) in western Kyūshū., Japan.

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H. pulchella UGA Voucher (Currie No. 100) ▲

Sobosan (祖母山) in situ ► Miyazaki-ken (宮崎県)

H. pycnophylla F. Maekawa ► J. Japanese Botany, 51, 3:80 1976 セトウチギボウシ = 瀬戸内 ギボウシ = Setouchi Gibōshi 西セトウチギボウシ = 西瀬戸内 ギボウシ = Nishi Setouchi Gibōshi Incorrect names: H. pycnophylla ‘Setouchi’ H. ‘Setouchi’ Urajiro Setouchi Gibōshi = 裏白 セトウチギボウシ AHS Name: H. pycnophylla Hab. Damp montane forests and along rocky river banks in Setouchi/Inland Sea Region (瀬戸内海); Ōshima Island (大島) on Genmeizan (源明山); also in Yashirojima (屋代島) on Monjusan (文珠山) Kanouzan (嘉納山) at higher elevations. Setouchi Gibōshi, named for Setouchi Region (瀬戸内海) and “pycnophylla” is its pycnos = close (many-petioled) leaf (phyll- = leaf) arrangement.

H. pycnophylla (in situ) (Notice Seedlings to Right)

AHS Trek ▪ Dr. R. Olson – Mt. Genmei (源明山), Ōshima Island (大島)

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■

H. pycnophylla Maekawa 1976 (in situ ▪ notice seedlings) Seen from below to shown white backs; Genmeizan (源明山)

Ōshima (大島); Ōshima-machi (大島町); Ōshima-gun (大島郡) Setouchi Region (瀬戸内海) ▪ ©AHS 1995/Olson

H. rectifolia Nakai (including in synonymy: Hosta atropurpurea; H. sachalinensis; H. apoiensis and the morph H. rectifolia f. albiflora (= H. ‘Shirobana Tachi’ = 白花立擬宝珠). Botanical Magazine, Tokyo, 44:26 and 58, Pl. 21 1930; and J. of the Faculty of Science, Imperial UniversityTokyo, Section 3 Botany, Vol. 5:410, 411, ic. 90

–93 940.

re)

1 H. re 沼)

HH No. 10068504 Hosta Hill R.G. © W.G. Schmid ▪ 6 Sept 2006

ctifolia (cult. ex Uryu Swamp (雨竜AHS Names: H. rectifolia var. rectifolia H. rectifolia (in horticultu

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Japanese Synonyms and Names:

(1940)

タチギボウシ = 立擬宝珠 = Tachi Gibōshi (1930) ウクr キナ = Ukurkina = Tachi Gibōshi (transliterated Ainu language = アイヌ語) エゾギボウシ = 蝦夷擬宝珠 = Ezo [also Yezo, Yeso] Gibōshi (G. Koidzumi) クロバナギボウシ = 黒花擬宝珠 = Kurobana Gibōshi (1930) 利尻ナギボウシ = 利尻擬宝珠 = Rishiri Gibōshi [from island Rishiri-tō (利尻島)] オゼギボウシ = 尾瀬擬宝珠= Oze Gibōshi [from Ozegahara Swamp 尾瀬ヶ原] オゼミズギボウシ =尾瀬水擬宝珠 Oze Mizu Gibōshi [from Ozegahara Swamp] 白花タチギボウシ = 白花立擬宝珠 = Shirobana Tachi Gibōshi (white-flowered) Hab.: H. rectifolia is a hygrophyte in sunny, wet meadows, and on shallow banks of lakes and rivers; volcanic plateaus with shallow lakes and ponds forming swamps like the Uryū Wetland (Uryū-numashitsugen; 雨竜沼湿原) located near Uryū-cho (雨竜町) in Sorachi-shichō (空知

支庁). Main habitat is in Hokkaidō (北海道) and the southern part of Sakhalin (Karafuto; 樺太). Sporadic in the southern Kurile Islands (Chishima Islands (千島列島) and the Russian Maritime Province of Primorsky Krai (Примо́рский край). Also in the Tōhoku region (Tōhoku-chihō; 東北地方), including Aomori, Akita and Iwate prefectures.

♠ H. rectifolia var. rectifolia f. pruinosa Maekawa J. of the Faculty of Science, Imperial University Tokyo, Sect. 3 Botany, Vol. 5:413 1940 トノコタチギボウシ = 砥の粉

立擬宝珠 = Tonoko Tachi GibōshiH. rectifolia ‘Pruinosa’ hort. incorrect. AHS Names: H. rectifolia var. rectifolia H. rectifolia f. pruinosa (Cultivated)

Kushiro-shichō (釧路支庁) (HH No.10568604) Hosta Hill R.G. ▪ W.G. Schmid ▪ 9 Aug 2006

f. pruinosa H. rectifolia f. pruinosa H. ’Tonoko Tachi’

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Hab.: Occasionally found in wetland habitats but more so in moist forest areas. It occurs in all parts of Hokkaidō (北海道) and the six northern prefectures of Tōhoku region (Tōhoku-chihō; 東北地方).

H. rectifolia f. pruinosa (Cultivated) Kushiro-shichō (釧路支庁) (HH No.10568604) Hosta Hill R.G. ▪ W.G. Schmid ▪ 13 July 2006

H. rectifolia var. australis W.G. Schmid The genus Hosta: Gibōshi Zoku (ギボウシ属). Timber Press; 1991 (in scheda: No. 6560 (643/225 Coll. No. 1217; M. Mitsushima); Uni. of Tokyo Herb. (東京大學綜合博物館標本館) マルバタチギボウシ = 円葉立擬宝珠 =

aruba Tachi Gibōshi (Schmid 1991)

alis . ‘Maruba Tachi’

ichō

Hokkaidō (北海道) [Court.:©3pro tok2 kasya]

M AHS Names: H. rectifolia var. austrH H. rectifolia var. australis (in situ) ► (Shiribeshi-shichō; 後志支庁); Shiribeshish(後志支庁), near Makubetsu-chō (幕別町)

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H. rectifolia var. australis (in situ) ► Aomori-ken (青森県); Hakkoda Mountains (八甲田山); Sukayu (酸ヶ湯), near Aomori City (青森市) ; ©W.G. Schmid 1989 ギボウ属 著者 Hab.: H. rectifolia var. australis is a hygrophyte similar to H. rectifolia forma typica. It is native further south in Tōhoku Tōhoku-chihō (東北地方), including Aomori, Akita and Iwate prefectures. Records indicate it grows in Gunma-ken ( 群馬県), Ozegahara moor (尾瀬ヶ原), Tone-gun (利根郡). In sunny, swampy meadows, on shallow banks of lakes and rivers, and volcanic plateaus with shallow ponds. Populations also in Hokkaidō (北海道) on Youtei-zan (羊蹄山) in Niseko-Shakotan-Otaru Kaigan Quasi-National Park (ニセコ積丹 小樽海岸国定公園).

♠ H. rohdeifolia f. viridis Maekawa J. of Japanese Botany, 13:897 ic. f. 3 1937 (pro parte); J. of the Faculty of Science, Imperial University Tokyo, Section 3 Botany, Vol. 5:406-407 1940. アオバオモトギボウシ = 青葉万年青 擬宝珠 = Aoba Omoto Gibōshi (Omoto stands for Rhodea japonica) AHS Name: H. rohdeifolia f. viridis H. rohdeifolia f. viridis (cult.) (ex Hieizan; 比叡山;Kyōtō-fu (京都府) Hosta Hill R.G. (V.: HH 1307/2004) © W.G. Schmid ▪ 21 April 2007

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Hab.: The all-green taxon H. rhodeifolia f. viridis species was found on Hiei-zan (比叡山) in Yamashiro-no kuni (山城国), near Sakamoto (坂本), Lake Biwa (琵琶

湖). It is rarely seen in cultivation. Its name Aoba Omoto stands for the leaf shape being similar to that of Rhodea japonica. Variegated forms formerly known under the incorrect name H. rhodeifolia `Aureomarginata' are now known under the Japanese name H. ‘Kifukurin Omoto’ (= 黄覆輪 万年青擬宝珠) ►►►

♠ H. rupifraga Nakai. Botanical Magazine, Tokyo, 44:27, ic. f. 3 1930; and Maekawa: Botanical Magazine, Tokyo, 52:42, ic. f. 2 1938. ハチジョウギボウシ = Hachijō Gibōshi 섬비비추 = Seom-bi-bi-shu (Kor.) AHS Name: H. rupifraga

H. rupifraga cult.▼ (in situ ►) H. rupifraga Hachijō-Fuji; Coll. J.E. Craig; Voucher CDS No. 436/1969;Hosta Hill R.G. HH 1988 © W.G. Schmid Coll. Mt. Nishi (八丈富士 on Hachijō-Fuji, 854m/2802 ft.) (Cult. specimen accessed at the NW-facing caldera rim 1969).

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Hab.: This taxon’s small, allopatric populations are limited to the rim of the Hachijō-Fuji caldera, at elevation 2,500 feet (845 meters) and just below. In spring it arises together with Astilbe hachijoensis and other alpines in full sun but by early summer the leaf mounds are overshadowed by tall grasses and stay in this shade for the remainder of the season. Although it has unique morphology, it is often confused with H. longipes var. latifolia. Fujita (1976), held this taxon to be synonymous with H. longipes var. latifolia, but RAPD analysis (Y. Yu, 2002; Sauve, R.J., S. Zhou, Y. Yu, and W.G. Schmid. 2005) shows that H. rupifraga and H. longipes var. latifolia are differentiated.

♠

H. shikokiana (in situ) ►▲ Higashiakaishiyama 東赤石山, Ehime-ken (愛媛県); high mountain district of (山地) Shikoku (四国). Court.: ©oNLINE 植物アルバム and 萩生の森 H. shikokiana N. Fujita Acta Phytotaxonomica et Geobotanica, Vol. 27, 3/4:93, 1976. シコクギボウシ = Shikoku Gibōshi = Shikoku Hosta AHS Name: H. shikokiana

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Hab: In open mountain valleys, on rocks and on outcrops along ridgelines of mountains in Ehime-ken (愛媛県) at 1300–1800 AMSL on Mt. Higashiakaishi (東赤石山), Mt. Akaishi (赤石岳), Mt. Nishiakaishi (西赤石山), Mt. Kanpu, Mt. Ishizuchi (石鎚山) and in Kōchi-ken (高知県) Shiraga-yama (白髪山), western Shikoku, Japan (from topotypes to show distribution).

♠ H. sieboldiana (Hooker) Engler In Engler and Prantl, Die Natürlichen Pflanzenfamilien, 2/5:39–40 1888. とうギボウシ = Tō Gibōshi

AHS Name: H. sieboldiana (= H. ‘Sieboldiana’) H. sieboldiana ►Author with historic voucher specimens cultivated at the Munich Botanic Garden (of Ludwig-Maximilians-Universität München-Nymphen-burg) © W.G. Schmid 1984 As cultivated in gardens today, H. sieboldiana is not of Japanese origin. It was first published by European botanists and it is the “European type” grown in Western gardens that is considered here as the “species” as described by Hylander (1954). The European type specimens are based on cultivated, hybridized sources. The taxon most often considered the “standard” form in Western gardens, namely H. sieboldiana ‘Elegans’, is known to be a hybrid made by Arends (1905) and named H. ‘Robusta’ (= H. ‘Tokudama’ × H. sieboldiana (a selected form of von Siebold’s importations). In 1984, I observed five of the Arends hybrids at the (historical) gardens of Fachhoch-schule (University) of Weihenstephan in Freising, FRG (see Type 1 photograph on page 16). The other types have similar macromorphology as shown and all look alike and are offered under the names H. sieboldiana or H. ‘Elegans’. In Japan, H. sieboldiana has been cultivated for centuries and includes many different phenotypes Maekawa [1940] named four, all originating in gardens. Based on research conducted by Hylander, Hensen and Schmid, the difference between H. montana and H. sieboldiana is finally established and gardeners can easily identify them:

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1) H. sieboldiana: Leaves larger, 18 in. by 11 in. (46 by 28 cm), with 16 to 19 vein pairs; yellow anthers (before dehiscing); the leaves are bluish grey, corrugated/dimpled and covered with bluish-gray indumentum both sides; the scapes are 20-36 inches (50 – 90 cm) tall, first erect, then bending (data for mature plants).

2) H. montana: Leaves smaller, 12.5 in. by 9.5 in. (32 by 24 cm), with 13 to 16 vein pairs deeply impressed on top; purple anthers (before dehiscing), the leaves are smooth dull green with no indumentum; the scapes are 50-60 inches (1.25-1.5 m) tall and erect (data for mature plants). As to whether this taxon should be considered a species is another matter. H. montana inhabits large parts of the Japanese Islands. H. sieboldiana as cultivated is present in many Japanese gardens. However, allopatric popula-tions have not been confirmed in the wild. Since the name has seen widespread use in horticulture the world over, keeping it as H. sieboldiana is a practical solution.

H. (sieboldiana) ◄ ‘Gray Cole’

(Ex Gray Cole. N.) Kuk’s Forest

Nursery (1985) © W.G. Schmid

1985

END PART 3

H. (sieboldiana) ‘Elegans’ (Type 1) One of 5 specimens obtained from K. Foerster (ex G. Arends) Fachhochschule (University) of Weihenstephan © W.G. Schmid August 1984

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